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Nactus grevifer ZUG, 2020

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Torricelli Slender-toed Gecko 
SynonymNactus grevifer ZUG 2020: 43 
DistributionPapua New Guinea (East Sepik Province, Kumnatei, Torricelli mountain range, Bom area of Madang)

Type locality: Papua New Guinea, West Sepik Province, Kumnatei [−3.45° 142.12°]  
Reproduction 
TypesHolotype. IRSNB 15802Q, adult male, collected by Benoit Mys in early 1980s.
Paratypes. IRSNB 15802A–F, adult females with the same collecting data as the holotype, and IRSNB 15802K–N, adult males with the same collecting data as the holotype. 
DiagnosisDefinition. A bisexual taxon of geckos (Gekkoninae) with moderate adults (♀♀ 47–57 mm, ♂♂ 47–61 mm SVL) with keeled subcaudals, no to small postmental scales (0–12, median 6), no or slight postmental-infralabial (0–2) contact, moderate number of dorsal tubercle rows (13–16), and mod- erately high number of tubercles (31–39) in parasagittal row (TubRow), strongly tuberculate dorsal surface of thigh and crus (large tubercles forming sheath-like surface), and modest num- ber of precloacal pores (8–11) in males (females without pores). Diagnostic summary in Table 8.

In life, coloration has not been reported; in alcohol it was not recorded.

Variation. Females and males from Kumnatei (local- ity 38, n = 17) display no striking metric differences (adult ♀♀ 54.7, 51.8–57.1 mm; adult ♂♂ 55.2, 51.8–61.1 mm SVL) and also share similar proportions (total Kumnatei adult sample): TrunkL/SVL 42.2%, 37%–49%; HeadL/SVL 26.3%, 25%– 29%; HeadW/SVL 18.1%, 16%–17%; HeadW/HeadL 69.3%, 64%–74%; EyeD/SVL 5.2%, 5%–6%; EyeD/HeadL 20.1%, 18%–23%; NarEye/HeadL 31.0%, 30%–32%; Interorb/HeadL 22.3%, 17%–28%; SnW/HeadL 13.4%, 11%–15%.
Typical head scale pattern of small, granular scales except for those bordering nares and mouth. Snout with large rostral scale bearing middorsal cleft posteriorly; large supralabials (3, 3–4) and infralabials (3, 2–4) in front of anterior edge of orbit, first supra- and infralabial largest of respective series. Chin with large trapezoidal mental and no to small postmentals (Postm = 4, 0–10, single specimen lacking postmental) and usually no genial scale between postmental and first surpalabial (0, 0–2 PmLab); dorsally, trunk with 14, 13–16 (DorsTub) enlarged tubercles transversely on dorsum and with 35, 33–39 tubercles (TubRow) longitudinally along trunk; 10, 8–11 tubercle rows transversely between hindlimbs (TubHip). Scattering of large tubercules on dorsal half of forelimb and sheath of tubercles on hindlimb (TubHindl = 3). Tail with small uniform scales dorsally and lat- erally, ventrally with uniform, small keeled scales (Subcaud = 0 with exception of one individual with smooth subcaudals). Usu- ally, 2 cloacal spurs (2, 1–3 CloacS) on each side. Ventral scales from chin to vent small, granular, somewhat larger on chest and abdomen. Precloacal pores, males 8–10, females 0. Forefoot with narrow digits, their lengths usually 3>4≈2>5≈1; 17, 16–19 lamel- lae beneath fourth digit (4FingLm), 6, 5–8 scale rows on palm; hindfoot with narrow digits, their lengths 4≈3>5>2>1; 22, 19–24 lamellae beneath fourth toe (4ToeLm).
A distant sample (locality = 21) from the Bom area contains individuals identified as N. allenallisoni (n = 2) and N. grevifer (n = 5). In spite of the distance from the grevifer type locality, I treat the latter specimens as that species. They are somewhat smaller (adult ♀♀ 54.7, 51.8–57.1 mm; adult ♂♂ 55.2, 51.8–61.1 mm SVL) and also share proportional differences (total Kumnatei adult sample): TrunkL/SVL 42.2%, 37%–49%; HeadL/SVL 26.3%, 25%–29%; HeadW/SVL 18.1%, 16%–17%; HeadW/HeadL 69.3%, 64%–74%; EyeD/SVL 5.2%, 5%–6%; EyeD/HeadL 20.1%, 18%–23%; NarEye/HeadL 31.0%, 30%–32%; Inter- orb/HeadL 22.3%, 17%–28%; SnW/HeadL 13.4%, 11%–15%. They also possess some differences in scalation. Large supralabi- als (3, 3–3) and infralabials (3, 2–3) in front of anterior edge of orbit, first supra- and infralabial largest of respective series. Chin with large trapezoidal mental and small to medium postmentals (Postm = 8, 6–12) and usually no genial scale between postmen- tal and first supralabial (0, 0–1 PmLab); dorsally, trunk with 14, 13–15 (DorsTub) enlarged tubercles transversely on dorsum and with 29, 25–30 tubercles (TubRow) longitudinally along trunk; 8, 7–8 tubercle rows transversely between hindlimbs (TubHip). Scattering of large tubercules on dorsal half of forelimb and sheath of tubercles on hindlimb (TubHindl = 3). Tail with small uniform scales dorsally and laterally, ventrally with uniform, small keeled scales (Subcaud = 0, with exception of one individual with smooth subcaudals). Usually, 2 cloacal spurs (2, 1–2 CloacS) on each side. Ventral scales from chin to vent small, granular, somewhat larger on chest and abdomen. Precloacal pores, females 0; the single male has 8 precloacal pores and a single femoral pore on each side. Forefoot with narrow digits, with 14, 13–15 lamellae beneath fourth digit (4FingLm), 5, 5–6 scale rows on palm; hindfoot with narrow digits, 18, 16–20 lamellae beneath fourth toe (4ToeLm). 
CommentDistribution: see map in Zug 2020: 47 (Fig. 18). 
EtymologySpecific name derives from the impres- sion of leg armor by the heavily tuberculate outer surface of the hindlimbs, hence wearing leg armor, that is, greaves. “Greave” derives from the Old French greve for leg armor. The Latin fero refers to “carrying,” and its suffix is ifer, hence grevifer, or wearer of leg armor. The specific name is a noun in apposition. 
References
  • Zug, George R. 2020. Diversity in Pacific Slender-Toed Geckos, Nactus pelagicus Complex (Reptilia: Squamata), of New Guinea and Adjacent Islands. Smithsonian Contributions to Zoology, (651): x + 92 pages - get paper here
 
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