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Nactus inundatus ZUG, 2020

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Fly River Slender-toed Gecko 
SynonymNactus inundatus ZUG 2020: 69 
DistributionPapua New Guinea (flood plain of the Fly River north to Lake Murray), likely Indonesia (Papua)

Type locality: Papua New Guinea, Western Province, Emeti [−7.8547° 143.2451°].  
Reproduction 
TypesHolotype. USNM 325059, adult male, collected by Fred Parker on 1 November 1971 (Figure 34).
Paratypes. Adult males, USNM 195757, 325063, and adult females, USNM 325061–062, 325066, with same collecting data as holotype. 
DiagnosisDefinition. A bisexual taxon of geckos (Gekkoni- nae) with moderate adults (♀♀ 48–58 mm, ♂♂ 45–54 mm SVL) with keeled subcaudals, postmental scales usually absent (0–8, median 0), moderate number of dorsal tubercle rows (11–15) and usually numerous tubercles (29–40, usually ≥32) in para- sagittal row (TubRow), no tubercles present on dorsal surface of thigh and crus, and precloacal pores in males (13, 9–15), females usually with pores (4, 0–13) but fewer than in males. Diagnostic summary in Table 8.

Variation. Females and males display nearly iden- tical metrics (adult ♀♀ 50.9, 48.0–58.1 mm; adult ♂♂ 50.7, 44.9–53.9 mm SVL) and share similar proportions (total adult sample): TrunkL/SVL 42.0%, 36%–47%; HeadL/SVL 26.9%, 25%–28%; HeadW/SVL 18.0%, 16%–20%; HeadW/HeadL 67.0%, 63%–72%; EyeD/SVL 6.9%, 6%–8%; EyeD/HeadL 25.7%, 23%–30%; NarEye/HeadL 30.8%, 26%–34%; Inter- orb/HeadL 27.0%, 23%–31%; SnW/HeadL 14.8%, 12%–17%. Typical head scale pattern of small, granular scales except for those bordering nares and mouth. Snout with large rostral scale with middorsal depression and cleft posteriorly; suprana- sals moderate sized, usually in contact on midline or separated by small granular scale (internasal); large supralabials (3, 3–4) and infralabials (3, 3–4) in front of anterior edge of orbit, first supra- and infralabial largest of respective series. Chin with large trapezoidal mental and usually (~85%) lacks postmen- tals (Postm = 0, 0–8); dorsally, trunk with 13, 11–15 (DorsTub) enlarged tubercles transversely on dorsum and with 36, 29–40 tubercles (TubRow) longitudinally along trunk; 8, 7–10 tubercle rows transversely between hindlimbs (TubHip). No enlarged tubercles on crus and femur (TubHindl = 0). Tail with small uni- form scales dorsally and laterally, ventrally with uniform, small keeled scales (Subcaud = 0). Usually, pair of cloacal spurs (2, 1–2 CloacS) on each side. Ventral scales from chin to vent small, granular, somewhat larger on chest and abdomen. Precloacal pores, males 9–15, females usually with pores 0–13. Forefoot with narrow digits, their lengths usually 3≈4>2≈5>1; 16, 13–18 lamellae beneath fourth digit (4FingLm), 6, 5–7 scale rows on palm; hindfoot with narrow digits, their lengths 4>3>5>2>1 or
3≈4>2≈5>1; 26, 18–22 lamellae beneath fourth toe (4ToeLm). Coloration in life not reported. In alcohol, color and pat- tern match the general description of the holotype. The dor- sal trunk pattern of transverse bars varies from faded to dark; in some dark bars divided at dorsal midline, from 5 to 6 bars between shoulders to sacrum. Forelimbs typically mottled dark and medium brown; hindlimbs (thigh) varying from mottled to
horizontal light bar posteriorly; venter uniform light brown. 
CommentDistribution: see map in Zug 2020: 39 (Fig. 12). 
EtymologyThe specific name derives the Latin inundatus in reference to the origin of the species’ landscape development from repeated flooding by the region’s rivers, espe- cially the Fly River. 
References
  • Zug, George R. 2020. Diversity in Pacific Slender-Toed Geckos, Nactus pelagicus Complex (Reptilia: Squamata), of New Guinea and Adjacent Islands. Smithsonian Contributions to Zoology, (651): x + 92 pages - get paper here
 
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