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Nactus kamiali ZUG, 2020

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Kamiali Slender-toed Gecko 
SynonymNactus kamiali ZUG 2020: 51 
DistributionPapua New Guinea (central Morobe Province [Kamiali] to Northern Province [Mount Lamington])

Type locality: Papua New Guinea, Morobe Province, Kamiali Wildlife Management Area, 1.3 km N and 6.2 km W of Cape Dinga, −7.296° 147.093°.  
Reproduction 
TypesHolotype. BPBM 25964, adult female, collected by Allen Allison on 5 September 2004 (Figure 21).
Paratypes. BPBM 25965, 25967–968, PNGM 17181, adult females, BPBM 25971, adult male with same collecting data as holotype; USNM 576310, adult male, CAS 249851, adult female, from Kamiali village, Kamiali Biological Research Station, 7°17′45.348′′S, 147°05′58.524′′E, Morobe Province, Papua New Guinea, collected by R. N. Fisher, J. Stanford, J. Richmond, and B. Iova on 22 May 2010. 
DiagnosisDefinition. A bisexual taxon of geckos (Gekko- ninae) with large adults (♀♀ 61–73 mm, ♂♂ 60–70 mm SVL) with keeled subcaudals, variable-sized postmental scales (2–19, usually ≥8), no or slight postmental-infralabial (PmLab = 0–1) contact, moderate number of dorsal tubercle rows (17–20) and moderate number of tubercles (27–34, usually ≥28) in parasagit- tal row (TubRow), tubercles on dorsal surface of thigh and crus, and precloacal pores (9, 8–11) in males, females lack pores (0). Diagnostic summary in Table 8.

Coloration in life not reported.

Variation. Females and males display slight metric differences (adult ♀♀ 67.3, 61.5–73.1 mm; adult ♂♂ 66.8, 59.5– 69.5 mm SVL), although sharing similar proportions (total adult sample): TrunkL/SVL 38.6%, 35%–45%; HeadL/SVL 27.8%, 27%–29%; HeadW/SVL 18.2%, 15%–20%; HeadW/HeadL 65.6%, 56%–71%; EyeD/SVL 6.7%, 6%–8%; EyeD/HeadL 24.1%, 21%–29%; NarEye/HeadL 33.2%, 30%–43%; Inter- orb/HeadL 25.2%, 22%–30%; SnW/HeadL 14.5%, 13%–16%.
Typical head scale pattern of small, granular scales except for those bordering nares and mouth. Snout with large rostral scale bearing middorsal cleft posteriorly; supranasals moderate sized, usually in contact on midline or separated by small granu- lar scale (internasal); large supralabials (4, 3–4) and infralabi- als (3.5, 3–4) in front of anterior edge of orbit, first supra- and infralabial largest of respective series. Chin with large trapezoi- dal mental and usually moderate-sized postmentals (Postm = 10, 2–19) and usually no genial scale between postmental and first supralabial (0, 0–1 PmLab); dorsally, trunk with 19, 17–20 (DorsTub) enlarged tubercles transversely on dorsum and with 30, 27–34 tubercles (TubRow) longitudinally along trunk; 10, 8–10 tubercle rows transversely between hindlimbs (TubHip). Uniform covering of small tuberculate-like scale on dorsal half of forelimb and enlarged tubercles on crus and femur (TubHindl = 2). Tail with small uniform scales dorsally and laterally, ventrally typically with uniform, small keeled scales (Subcaud = 0, 0–1). Usually, single cloacal spurs (1, 1–2 CloacS) on each side. Ventral scales from chin to vent small, granular, somewhat larger on chest and abdomen. Precloacal pores, males 8–11, females 0. Forefoot with narrow digits, their lengths usually 3≈4>2>5>1; 18, 16–19 lamellae beneath fourth digit (4FingLm), 5, 4–7 scale rows on palm; hindfoot with narrow digits, their lengths 4≈3>2≈5>1; 22, 20–23 lamellae beneath fourth toe (4ToeLm).

Coloration in life, dorsal and lateral background variable light to medium brown with distinct to diffuse dark transverse bars middorsally continuous or separated, usually dark nape bar and always mid-cervical bar, 5 bars from shoulder to trunk; dark-brown sacral and first caudal bars always evident; top of head unicolor to scattered dark marks. Dorsally, limbs usually dark brown mottled, posterior thighs of most individuals with pair of dark horizontal stripes enclosing near-white stripe. In alcohol, background tan to medium brown dorsally and later- ally. Top of head with medial darker-brown figure from interor- bital onto snout, supralabials large, brown with white to cream posterior margin, infralabials cream to dusky white. Nearly uni- form tan dorsally from crown onto base of tail. Venter uniform light tan from chin onto tail. 
CommentDistribution: see map in Zug 2020: 53 (Fig. 22). 
EtymologyThe specific name derives directly from the geographic place name, Kamiali, where the first specimens were discovered by Allen Allison. 
References
  • Zug, George R. 2020. Diversity in Pacific Slender-Toed Geckos, Nactus pelagicus Complex (Reptilia: Squamata), of New Guinea and Adjacent Islands. Smithsonian Contributions to Zoology, (651): x + 92 pages - get paper here
 
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