You are here » home › advanced search › search results › Naja nigricollis

Naja nigricollis REINHARDT, 1843

iNaturalist logo

Can you confirm these amateur observations of Naja nigricollis?

Add your own observation of
Naja nigricollis »

Find more photos by Google images search: Google images

Higher TaxaElapidae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Black-necked Spitting Cobra
G: Afrikanische Speikobra 
SynonymNaja nigricollis REINHARDT 1843: 369
Naja nigricollis var. crawshayi GÜNTHER 1893
Naja nigricillis var. occidentalis BOCAGE 1895
Naja nigricollis — SCHMIDT 1923: 128 (fide BROADLEY 1968)
Naja nigricollis atriceps LAURENT 1955: 135
Naja nigricollis crawshayi — LAURENT 1956: 298
Naja nigricollis occidentalis — LAURENT 1973
Naja nigricollis nigricollis — BRANCH 1981: 29
Naja nigricollis nigricollis — BROADLEY & HOWELL 1991: 24
Naja nigricollis nigricollis — WELCH 1994: 93
Naja crawshayi — BROADLEY & COTTERILL 2004
Naja nigricollis — CHIRIO & INEICH 2006
Naja crawshayi — BRANCH 2008
Naja nigricollis — WALLACH et al. 2009
Afronaja nigricollis — WALLACH et al. 2014: 10
Afronaja crawshayi — WALLACH et al. 2014: 9
Naja (Afronaja) nigricollis — CONRADIE et al. 2016
Naja nigricollis — SPAWLS et al. 2018: 555 
DistributionAngola, Benin, Burkina Faso, Cameroon, Central African Republic, Chad, Democratic Republic of the Congo (Zaire, except in the center), Congo, Ethiopia, Gabon, Ghana, Guinea-Bissau, Guinea (Conakry), Ivory Coast, Kenya, Liberia, Mali, Niger, Nigeria, Senegal, Sierra Leone, Gambia (HÅKANSSON 1981), Mauritania, Sudan (Jumhūriyyat), Republic of South Sudan (RSS), Tanzania, Somalia, Togo, Uganda, Zambia

nigricollis: Savannas from Senegal east to Kenya and south to Angola, Zambia, and Tanzania; Type locality: Guinea = coast of Ghana.

atriceps (invalid): Type locality: Mugera, Ruyigi, Burundi.

crawshayi: Democratic Republic of the Congo (Katanga), Congo (Brazzaville) through N Malawi to Tanzania; Type locality: Lake Mweru [= Lake Meru, N Zambia]  
Reproductionoviparous 
TypesHolotype: ZMUC 65371
Holotype: BMNH 1946.1.21.69, 1946.1.23.80, skin of an adult [crawshayi]
Holotype: RMCA (= MRAC = RGMC) 17529. Paratypes: ZFMK 50004 (ex-IRSNB 2.068), IRSNB 2.069 - 2.081 [atriceps] 
DiagnosisDiagnosis (subgenus Afronaja): Intracranial (dorsal) anterior Vidian canal position, 2–3 solid maxillary teeth, six supralabials (except some N. nubiae and N. pallida) with penultimate (fifth or sixth) shield low, combination of two preoculars and two or three anterior temporals, rostral as broad as deep; internasals as long as prefrontals, dorsal scales matte or moderately shiny, and fangs adapted for spitting.


Additional details (445 characters) are available for collaborators and contributors. Please contact us for details. 
CommentVenomous!

Subspecies: BOYCOTT & HAACKE (1979) noted intergradation between nigricincta and woodi in Southern Namibia. Naja nigricollis nigricincta BOGERT 1940 has been elevated to full species status with N. n. woodi as subspecies.

Type species: Naja nigricollis REINHARDT 1843 is the type species of the subgenus Afronaja WALLACH, WÜSTER & BROADLEY 2009. Note that Raymond Hoser had named the same group of cobras “Spracklandus” 6 months earlier. Rhodin et al. 2015 requested to declare the Australasian Journal of Herpetology (AJH) Volumes 1–24 unavailable for nomenclatural purposes, thereby rendering the name Spracklandus unavailable. ICZN Opinion 2468 (2021) declined to confirm that Spracklandus Hoser is an available name and thus effectively left the case open to the community to decide. See also the comment by Krell 2021.

Synonymy: Kaiser et al. 2013 considered the generic name Spracklandus Hoser 2009 invalid and rejected its use instead of Naja (Afronaja). Hoser (2013) requested to retain Spracklandus in ICZN Case 3601. Naja nigricollis var. crawshayi GÜNTHER 1893 has been removed from the synonym of N. nigricollis. According to Laurent (1956: 298; 1964: 120): the crawshayi of Katanga are different from nigricollis in ventral counts and head color (brown in adults, appears to be also the case in younger nigricollis …). Laurent 1964: 120 discusses the difference between crawshayi and occidentalis (from Angola) and the specimens from Broadley & Pitman (1960) attributed to crawshayi, which he attributes to occidentalis or possibly atriceps. W. Wüster recommended to treat N. crawshayi as synonym of nigricollis (pers. comm., 29 Sep 2016).

Distribution: see map in BROADLEY 1974.

Diet: bird eggs, among other things (Bob Golding 2020, online). 
EtymologyNamed after the black coloration of the neck, from Latin niger, nigra, nigrum = black, and Latin collis = neck. 
References
  • Abraham, G. 1981. Interessante Jugendfärbung bei Naja nigricollis. Sauria 3 (2): 21-21 - get paper here
  • Auerbach, R.D. 1987. The Amphibians and Reptiles of Botswana. Mokwepa Consultants, Botswana, 295 pp.
  • Baptista NL, António T, Branch WR. 2019. The herpetofauna of Bicuar National Park and surroundings, southwestern Angola: a preliminary checklist. Amphibian & Reptile Conservation 13(2) [Special Section]: 96–130 (e203) - get paper here
  • Barnett, Linda K. 2001. The herpetofauna of Abuko Nature Reserve, the Gambia. Herpetological Bulletin (77): 5-1 - get paper here
  • Barnett, Linda K. & Emms, Craig 2005. Common reptiles of The Gambia. Rare Repro, Hailsham, East Sussex, 24 pp.
  • Barts, Mirko 2018. Ein kleiner Reiseführer zu den Reptilien und Amphibien Namibias. Draco 16 (64): 6-23 [2015] - get paper here
  • Bauer, A.M., and Branch, W.R. 2003. The herpetofauna of the Richtersveld National Park, Northern Cape Province, Republic of South Africa. Herpetological Natural History 8: 111-160 [2001]
  • Bauer, Aaron M.; Branch, William R. & Haacke, Wulf D. 1993. The herpetofauna of the Kamanjab area and adjacent Damaraland, Namibia. Madoqua (Windhoek) 18 (2): 117-145. - get paper here
  • Bocage,J.V. du B. 1866. Lista dos reptis das possessões portuguezas d'Africa occidental que existem no Museu Lisboa. Jorn. Sci. Math. Phys. Nat. Lisboa 1: 37-56 - get paper here
  • Böhme, Wolfgang 2014. Herpetology in Bonn. Mertensiella 21. vi + 256 pp. - get paper here
  • Böhme, Wolfgang, Mark-Oliver Rödel, Christian Brede & Philipp Wagner 2011. The reptiles (Testudines, Squamata, Crocodylia) of the forested southeast of the Republic Guinea (Guinée forestière), with a country-wide checklist. Bonn zoological Bulletin 60 (1): 35-61 - get paper here
  • Boulenger, G. A. 1895. On the reptiles and batrachians obtained by Mr. E. Lort-Phillips in Somaliland. Ann. Mag. nat. Hist. (6) 16: 165-169 - get paper here
  • Boycott, R. C., and W. D. Haacke 1979. Note on the type-locality, distribution and juvenile coloration of Naja nigricollis woodi (Serpentes: Elapidae) and an account of the colour-pattern variation in intergrade populations. Annals of the Cape Provincial Museum (Nat. Hist.) 13: 31-36
  • Böhme, Wolfgang & Jeffrey Heath 2018. Amphibian and reptilian records from south-central Mali and western Burkina Faso. Bonn zoological Bulletin 67 (1): 59–69 - get paper here
  • Branch, W. R. 1981. The venomous snakes of southern Africa 2. Elapidae and Hydrophidae. Bull. Maryland Herp. Soc. 17 (1): 1-47 - get paper here
  • Branch, W.R. 2008. PROPOSED KALUKUNDI MINE (DEMOCRATIC REPUBLIC OF THE CONGO) - TERRESTRIAL FAUNA. ENVIRONMENTAL IMPACT ASSESSMENT REPORT (FINAL REPORT). A specialist report for Envirolution Consulting (Pty) Ltd., 83 pp.
  • Broadley, D. G. 1974. A review of the cobras of the Naja nigricollis complex in southwestern Africa (Serpentes: Elapidae). Cimbebasia (A) 2: 155-162
  • Broadley, D. G. & HOWELL, K. M. 1991. A check list of the reptiles of Tanzania, with synoptic keys. Syntarsus 1: 1—70
  • Broadley, D.G. 1958. The races of Naja nigricollis occurring in Rhodesia. J. Herp. Assoc. Rhodesia 2: 5–6 - get paper here
  • Broadley, D.G. 1968. A review of the African cobras of the genus Naja (Serpentes: Elapinae). Arnoldia (Rhodesia) (ser. 3) 29: 1-14
  • Broadley, D.G. 1991. The Herpetofauna of Northern Mwinilunga Distr., Northw. Zambia. Arnoldia Zimbabwe 9 (37): 519-538
  • Broadley, D.G.; Doria, C.T. & Wigge, J. 2003. Snakes of Zambia. An Atlas and Field Guide. Edition Chimaira, Frankfurt, 280 pp. [review in Sauria 26 (3): 21]
  • Broadley, Donald G. and F. P. D. Cotterill. 2004. The reptiles of southeast Katanga, an overlooked 'hot spot'. [Congo]. African Journal of Herpetology 53 (1): 35-61. - get paper here
  • Chifundera, K. 1990. Snakes of Zaire and their bites. Afr. Stud. Monogr. (Kyoto) 10(3): 137-157.
  • Chippaux, Jean-Philippe & Kate Jackson 2019. Snakes of Central and Western Africa. Johns Hopkins University Press, 448 pp. [detaileld review in HR 51 (1): 161] - get paper here
  • Chirio, L. 2009. Inventaire des reptiles de la région de la Réserve de Biosphère Transfrontalière du W (Niger/Bénin/Burkina Faso: Afrique de l’Ouest). [Herpetological survey of the W Transfrontier Biosphere Reserve area (Niger/Benin/Burkina Faso: West Africa]. Bull. Soc. Herp. France (132): 13-41 - get paper here
  • Chirio, L. & Lebreton, M. 2007. Atlas des reptiles du Cameroun. MNHN, IRD, Paris 688 pp.
  • Chirio, Laurent and Ivan Ineich 2006. Biogeography of the reptiles of the Central African Republic. African Journal of Herpetology 55(1):23-59. - get paper here
  • Cimatti, E. 2007. Namibia - Introduction to a vast territory. Reptilia (GB) (50): 58-67 - get paper here
  • Conradie W and Branch WR. 2016. The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation 10 (2) [Special Section]: 6–36 - get paper here
  • CONRADIE, WERNER; NINDA L. BAPTISTA, LUKE VERBURGT, CHAD KEATES, JAMES HARVEY, TIMOTEO JULIO & GOTZ NEEF. 2021. Contributions to the herpetofauna of the Angolan Okavango-Cuando-Zambezi river drainages. Part 1: Serpentes (snakes). Amphibian & Reptile Conservation 15(2): 244–278. - get paper here
  • Dobiey, M. & Vogel, G. 2007. Venomous Snakes of Africa - Giftschlangen Afrikas. Edition Chimaira, Terralog 15, 150 pp. - get paper here
  • Haagner, G.V.; Branch, W.R. 1993. Geographical Distribution - Naja nigricollis woodi. J. Herp. Assoc. Africa (42): 43-43 - get paper here
  • Haagner, G.V.; Carpenter, G. 1992. Notes on the Captilve Breeding of the Black Spitting Cobra, Naja nigricollis woodi. J. Herp. Assoc. Africa (41): 22-24 - get paper here
  • Haagner,G.V.; Branch,W.R. & Haagner,A.J.F. 2000. Notes on a collection of reptiles from Zambia and adjacent areas of the Democratic Republic of the Congo. Annals of the Eastern Cape Museum 1: 1 – 25
  • Hawkeswood, T.J. 2021. Time to end taxonomic vandalism by Wolfgang Wuster et al.: The Snakeman, Raymond Hoser’s publications are validly published and his names available according to the ICZN: Objective investigation finds Hoser’s taxonomic works as scientific best practi Calodema, 860: 1-51 - get paper here
  • Hayes, W.K., S.S. Herbert, J.R. Harrison & K.L. Wiley 2008. Spitting versus Biting: Differential Venom Gland Contraction Regulates Venom Expenditure in the Black-Necked Spitting Cobra, Naja nigricollis nigricollis Journal of Herpetology 42 (3): 453-460. - get paper here
  • Hellmich, W. 1957. Herpetologische Ergebnisse einer Forschungsreise in Angola. Veröff. Zool. Staatssammlung München 5: 1-91 - get paper here
  • Herrmann, Hans-Werner; Schmitz, Andreas; Herrmann, Patricia A. & Böhme, Wolfgang 2007. Amphibians and reptiles of the Tchabal Mbabo mountains, Adamaoua plateau, Cameroon. Bonner zoologische Beiträge 55 (1/2): 27-35 - get paper here
  • Hoser, R. 2009. A reclassification of the True Cobras; species formerly referred to the genera Naja, Boulengerina and Paranaja. Australasian J. Herpetol. 7: 1-15 - get paper here
  • Hoser, R.T. 2013. Case 3601 Spracklandus Hoser, 2009 (Reptilia, Serpentes, ELAPIDAE): request for confirmation of the availability of the generic name and for the nomenclatural validation of the journal in which it was published. Bull. Zool. Nomenclature 70 (4): 234-237 - get paper here
  • Hörold, Ralf & Heidrun 2018. Die Königspythons daheim b(g)esucht, Teil 2. Ophidia 12 (1): 18-25
  • Hughes, B. 2013. Snakes of Bénin, West Africa. Bull. Soc. Herp. France 144: 101-159
  • Hughes, D. F., et al. 2017. Geographic Distribution: Naja nigricollis (Black-necked Spitting Cobra). Herpetological Review 48 (2): 392 - get paper here
  • ICZN 2021. Opinion 2468 (Case 3601) – Spracklandus Hoser, 2009 (Reptilia, Serpentes, ElapidaE) and Australasian Journal of Herpetology issues 1–24: confirmation of availability declined; Appendix A (Code of Ethics): not adopted as a formal criterion for ruling Bulletin of Zoological Nomenclature 78: 42-45 - get paper here
  • Kaiser, H.; Crother, B.I.; Kelly, C.M.R.; Luiselli, L.; O’Shea, M.; Ota, H.; Passos, P.; Schleip, W.D. & Wüster, W. 2013. Best Practices: In the 21st Century, Taxonomic Decisions in Herpetology are Acceptable Only When Supported by a Body of Evidence and Published via Peer-Review. Herpetological Review 44 (1): 8-23
  • Krell, F. T. 2021. Suppressing works of contemporary authors using the Code's publication requirements is neither easy nor advisable. The Bulletin of Zoological Nomenclature, 78(1), 61-67 - get paper here
  • Lanza, B. 1983. A list of the Somali amphibians and reptiles. MONITORE ZOOLOGICO ITALIANO, new Ser., SUPPL. 18 (8): 193-247 - get paper here
  • Lanza, B. 1990. Amphibians and reptiles of the Somali Democratic Republic: check list and biogeography. Biogeographia, 14: 407-465 [1988] - get paper here
  • Largen, M.J.; Spawls, S. 2010. Amphibians and Reptiles of Ethiopia and Eritrea. Edition Chimaira, Frankfurt, 694 pp.
  • Largen,M.J. & Rasmussen,J.B. 1993. Catalogue of the snakes of Ethiopia (Reptilia Serpentes), including identification keys. Tropical Zoology 6: 313-434 - get paper here
  • Laurent, R.F. 1955. Diagnoses preliminaires des quelques Serpents venimeux. Rev. Zool. Bot. Afr., 51: 127-139
  • Loveridge, A. 1936. African reptiles and amphibians in the Field Museum of Natural History. Zool. Ser. Field Mus. Nat. Hist., Chicago, 22 (1): 1-122 - get paper here
  • Loveridge, A. 1956. On snakes collected in the Anglo-Egyptian Sudan by J.S. Owen, Esq. Sudan Notes Rec. 36: 37-56 [1955]
  • Loveridge, A. 1957. On a third collection of reptiles taken in Tanganyika by C.J.P. Ionides, Esq. Tanganyika Notes and Records 43: 1-19
  • Loveridge, Arthur 1929. East African reptiles and amphibians in the United States National Museum. Bull. US Natl. Mus. (151): 1-135 - get paper here
  • MADSEN, THOMAS & WOLFGANG WÜSTER. 2022. Black-necked spitting cobra Naja nigricollis Reinhardt, 1843 repels a large pack of banded mongooses (Mungos mungo). African Herp News 80: 75–78.
  • Marques, Mariana P.; Luis M. P. Ceríaco , David C. Blackburn , and Aaron M. Bauer 2018. Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola -- Atlas of Historical and Bibliographic Records (1840–2017). Proc. Cal. Acad. Sci. (Ser. 4) 65: 1-501 (Supplement II)
  • Menzies, J.I. 1966. The snakes of Sierra Leone. Copeia 1966 (2): 169-179. - get paper here
  • Mitchell, B.L. 1950. Some reptiles and amphibians of Nyasaland. The Nyasaland Journal 3 (2): 46-57 - get paper here
  • Monasterio, Camila 2016. The herpetofauna of the Dindefelo Natural Community Reserve, Senegal. Herpetology Notes 9: 1-6 - get paper here
  • Nneji, Lotanna Micah; Adeniyi Charles Adeola, , Agboola Okeyoyin, Ojo Cornelius Oladipo, Yohanna Saidu, Dinatu Samuel, John Yinamga Usongo, Babatunde Emmanuel Adedeji, Olatunde Omotoso, Akindele OluwatosinAdeyi, ObihAlexUgwumba,andAdiahaAldaAlexUgwum 2019. Diversity and distribution of amphibians and reptiles in Gashaka Gumti National Park, Nigeria. Herpetology Notes 12: 543-559 - get paper here
  • Padial, J. M. 2006. COMMENTED DISTRIBUTIONAL LIST OF THE REPTILES OF MAURITANIA (WEST AFRICA). Graellsia, 62(2): 159-178 - get paper here
  • Pauwels OSG, Morelle S, Albert J-L, Carlino P, Rahola N, Trape J-F. 2019. New reptile records from Lékédi Park and Haut-Ogooué Province, southeastern Gabon. Amphibian & Reptile Conservation 13 (1) [General Section]: 143–161 (e174) - get paper here
  • Pauwels, O.S.G. & Vande Weghe, J.P. 2008. Les reptiles du Gabon. Smithsonian Institution, Washington: 272 pp. - get paper here
  • Pietersen DW, Pietersen EW, Conradie W. 2017. Preliminary herpetological survey of Ngonye Falls and surrounding regions in south-western Zambia. Amphibian & Reptile Conservation 11(1) [Special Section]: 24–43 (e148 - get paper here
  • Pietersen, Darren, Verburgt, Luke & Davies, John 2021. Snakes and other reptiles of Zambia and Malawi. Struik Nature / Penguin Random House South Africa, 376 pp., ISBN 9781775847373
  • Pitman,C.R.S. 1974. A guide to the snakes of Uganda. Codicote, Wheldon & Wesley, L., 290 pp.
  • Pringle, JA. 1955. A new species of the spitting Cobra Naja nigricollis Reinhardt from the Cape Province. Ann. Natal Mus. 13: 253-254.
  • Rasmussen, J.B. & B. Hughes 1996. Description of some new snake species. I. [English translation of the original Danish text of T. Reinhardt 1843]. Steenstrupia 22: 13-39
  • Reinhardt, J. T. 1843. Beskrivelse af nogle nye Slangearter. Danske Vidensk. Selsk. Afhandl. 10: 233-279. - get paper here
  • Rhodin, A.G.J., H. Kaiser, P.P. van Dijk, W. Wüster, M. O’Shea, M. Archer, M. Auliya, L. Boitani, R. Bour, V. Clausnitzer, T. Contreras-MacBeath, B.I. Crother, J.M. Daza, C.A. Driscoll, O. Flores-Villela, J. Frazier, U. Fritz, A. Gardner, C. Gascon, 2015. Comment on Spracklandus Hoser, 2009 (Reptilia, Serpentes, ELAPIDAE): request for confirmation of the availability of the generic name and for the nomenclatural validation of the journal in which it was published (Case 3601; see BZN 70: 234–237; 71: 3 Bull. Zool. Nomenclature 72 (1):65-78
  • Robertson, I. A. D., B. M. Chapman and N. F. Chapman 1963. Notes on some reptiles collected in the Rukwa Valley, S. W. Tanganyika. Ann. Mag. Nat. Hist. Ser. 13, 5(55): 421‑432 - get paper here
  • Rödel,M.O.; Kouadio,K, & Mahsberg,D. 1999. Die Schlangenfauna des Comoé-Nationalparks, Elfenbeinküste: Ergänzungen und Ausblick. Salamandra 35 (3): 165-180 - get paper here
  • Schmidt, K. P. 1923. Contributions to the herpetology of the Belgian Congo based on the collection of the American Museum Congo Expedition, 1909-1915. Part II. Snakes, with field notes by Herbert Lang and James P. Chapin. Bull. Amer. Mus. nat. Hist. 49 (1): 1-146 - get paper here
  • Segniagbeto GLazcano. H., Trape J. F., David P., Ohler A., Dubois A. & Glitho I. A. 2011. The snake fauna of Togo: systematics, distribution and biogeography, with remarks on selected taxonomic problems. Zoosystema 33 (3): 325-360. DOI: 10.5252/z2011n3a4 - get paper here
  • Segniagbeto, G.H.; Dekawole, J.K.; Ketoh, G.K.; Dendi, D.; Luiselli, L. 2022. Herpetofaunal Diversity in a Dahomey Gap Savannah of Togo (West Africa): Effects of Seasons on the Populations of Amphibians and Reptiles. Diversity 14: 964
  • Senter, P. J., & Chippaux, J. P. 2022. Biogeography of snakes in Liberia: Review and synthesis of current knowledge. Ghana Journal of Science, 63(1), 29-62 - get paper here
  • Sow, Andack Saad; Duarte Vasconcelos Gonçalves, Fábio Vieira Sousa, Fernando Martínez- Freiría, Frederico Santarém, Guillermo Velo-Antón, Hamidou Dieng, João Carlos Campos, Saer Khayar Diagne, Zbyszek Boratyński, José Carlos Brito 2017. Atlas of the distribution of amphibians and reptiles in the Diawling National Park, Mauritania. Basic and Applied Herpetology - get paper here
  • Spawls, S., Branch, B. 1995. The Dangerous Snakes of Africa. Blandford, London, 192 pp.
  • Spawls, S.; Howell, K.; Drewes, R.C. & Ashe, J. 2002. A field guide to the reptiles of East Africa. Academic Press, 543 pp. [reviews in HR 34: 396 and Afr. J. Herp. 51; 147] - get paper here
  • Spawls, Stephen; Tomáš Mazuch & Abubakr Mohammad 2023. Handbook of Amphibians and Reptiles of North-east Africa. Bloomsbury, 640 pp. - get paper here
  • Spawls, Steve; Kim Howell, Harald Hinkel, Michele Menegon 2018. Field Guide to East African Reptiles. Bloomsbury, 624 pp. - get paper here
  • Sternfeld, R. 1910. Zur Schlangenfauna Deutsch-Südwestafrikas. Mehrere Fälle von Mimikry bei afrikanischen Schlangen. Mitt. zool. Mus. Berlin, 5: 51-60 - get paper here
  • Trape J-F and Mané Y. 2015. The snakes of Niger. Amphibian & Reptile Conservation 9(2) [Special Section]: 39–55 (e110) - get paper here
  • Trape, J.-F. 2023. Guide des serpents d’Afrique occidentale, centrale et d’Afrique du Nord. IRD Éditions, Marseille, 896 pp.
  • Trape, J.-F. & Mané, Y. 2004. Les serpents des environs de Bandafassi (Sénégal oriental). Bull. Soc. Herp. France 109: 5-34 - get paper here
  • Trape, J.-F. & Mané, Y. 2006. Guide des serpents d’Afrique occidentale. Savane et désert. [Senegal, Gambia, Mauritania, Mali, Burkina Faso, Niger]. IRD Editions, Paris, 226 pp. - get paper here
  • Trape, J.F. & R. ROUX-ESTÈVE 1995. Les serpents du Congo: liste commentée et clé de détermination. Journal of African Zoology 109 (1): 31-50
  • TRAPE, Jean-François & Marcel COLLET 2021. Nouvelles données sur les serpents du sud-est du Katanga (République démocratique du Congo). Bull. Soc. Herp. Fr. (2021) 179 | 11-26 - get paper here
  • TRAPE, JEAN-FRANÇOIS & CELLOU BALDÉ 2014. A checklist of the snake fauna of Guinea, with taxonomic changes in the genera Philothamnus and Dipsadoboa (Colubridae) and a comparison with the snake fauna of some other West African countries. Zootaxa 3900 (3): 301–338 - get paper here
  • Trape, Jean-François; Israël Demba Kodindo, Ali Sougoudi Djiddi, Joseph Mad-Toïngué & Clément Hinzoumbé Kerah 2020. The snakes of Chad: results of a field survey and annotated country-wide checklist. Bonn zoological Bulletin 69 (2): 367–393 - get paper here
  • Trape, Jean-François; Mané, Youssouph 2000. Les serpents des environs de Dielmo (Sine-Saloum, Sénégal). Bull. Soc. Herp. France 95: 19-35 - get paper here
  • Trape, Jean-François & Youssouph Mané 2017. The snakes of Mali. Bonn zoological Bulletin 66 (2): 107–133 - get paper here
  • Ullenbruch, K.; Grell, O.; Böhme, W. 2010. Reptiles from southern Benin, West Africa, with the description of a new Hemidactylus (Gekkonidae), and a country-wide checklist. Bonn Zool. Bull. 57 (1): 31-54 - get paper here
  • Wallach, V.; Wüster, W. & Broadley, D.G. 2009. In praise of subgenera: taxonomic status of cobras of the genus Naja Laurenti (Serpentes: Elapidae). Zootaxa 2236: 26–36 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Winchell, S. 2011. Kobras. Reptilia (Münster) 16 (89): 16-22 - get paper here
  • Wüster, W. & Broadley, D.G. 2007. Get an eyeful of this: a new species of giant spitting cobra from eastern and north-eastern Africa (Squamata: Serpentes: Elapidae: Naja). Zootaxa 1532: 51–68 - get paper here
  • Wüster, Wolfgang; Steven Crookes, Ivan Ineich, Youssouph Mané, Catharine E. Pook, Jean-Francois Trape, Donald G. Broadley 2007. The phylogeny of cobras inferred from mitochondrial DNA sequences: Evolution of venom spitting and the phylogeography of the African spitting cobras (Serpentes: Elapidae: Naja nigricollis complex). Molecular Phylogenetics and Evolution 45: 437–453 - get paper here
 
External links  
Is it interesting? Share with others:

As link to this species use URL address:

https://reptile-database.reptarium.cz/species?genus=Naja&species=nigricollis

without field 'search_param'. Field 'search_param' is used for browsing search result.



Please submit feedback about this entry to the curator