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Nucras holubi (STEINDACHNER, 1882)

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Higher TaxaLacertidae, Eremiadinae, Sauria, Lacertoidea, Squamata (lizards)
Subspecies 
Common NamesE: Holub’s Sandveld Lizard 
SynonymEremias holubi STEINDACHNER 1882
Nucras intertexta holubi — BOULENGER 1920: 20
Nucras intertexta holubi FITZSIMONS 1943: 320 (not of STEINDACHNER, fide BOYCOTT 1992)
Nucras holubi — JACOBSEN 1989
Nucras taeniolata holubi — BAUER et al. 1995
Nucras holubi — CONRADIE et al. 2011
Nucras holubi — EDWARDS et al. 2013
Nucras holubi — TELFORD et al 2022 
DistributionSouthern Africa, excluding the south-western and western Cape Province, Little and Great Namaqualand.

Recorded localities: Palapye Rd.; Serowe; Leeuwdoornsstad; Nylstroom; Doornkop, nr. Belfast; Dientje, nr. Vaalhoek; Newington; Ingwavuma; Makumbi; Empandeni; Rusape; Filabusi-Shabani; Monte Cassino, TIL Macheke (A.M.). Marandellas (A.M. &. K.M.). Kimberley (A.M., K.M., S A M . & T.M.). Hluhluwe (A.M. & N.M.). Potchefstroom (A.M. & T.M.). Umsinene Drift; nt. Inyamanzi (B.M.). Delport's Hope; Riverton; Belmont; Mafeking; Modder River; Jacobsdal; Taungs; Honeynestkloof; 'Witput; Warrendale; Lobatsi; Umvoma;: Ottoshoop; Premier Mine; Francistown; Molepoiole; Plumtroe (K.M.). Bulawayo (K.M., R.M. & S.A.M.). Junction Umfolosi Rivs. (N.M.). Mariepskop (P.E.M. & T.M.). Outjo; Kaoko Otavi; Otjitondua; Burghersdorp; Smithfield; Machudi; Junction Marico & Crocodile Rivs.; Bindura (SAM.). Acornhoek; Barberton (SAM. & T.M.). Odendaalsrust; Pretoria; Rustenburg;' Krabbefontein; Shilowane;: Pietersburg; Leydsdorp; Haenertsburg; Hectorspruit; Metlepetsi River; Blaauwberg; Silwane; Gravelotte; Medingen.; Perkoe, Olifants,' Riv. Dist.; btwn. Lake Fundusi & Entabeni; Punda Maria; Uanetsi Riv., E. Tvl.; Weenen; ,Mt. Silinda; Birchenough Bridge (T.M.) [from FITZSIMONS 1943: 322].  
Reproduction 
TypesLectotype: NMW 19056, designated by BROADLEY (1972); erroneously reported to be in the ZMB collection. 
Diagnosis 
CommentNucras taeniolata has been split into 3 species by Jacobsen (1989): Nucras taeniolata, N. holubi STEINDACHNER (1882) and N. ornata GRAY, J. E. (1864). 
Etymologynamed after the collector, E. Holub. 
References
  • Auerbach, R.D. 1987. The Amphibians and Reptiles of Botswana. Mokwepa Consultants, Botswana, 295 pp.
  • Bates, M.F.; Branch, W.R., Bauer, A.M.; Burger, M., Marais, J.; Alexander, G.J. & de Villliers, M.S. (eds.) 2014. Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, 512 pp.
  • Bauer AM, Murdoch M, Childers JL. 2020. A reevaluation of records of Sandveld lizards, Nucras Gray, 1838 (Squamata: Lacertidae), from northern Namibia. Amphibian & Reptile Conservation 14(3) [Taxonomy Section]: 231–250 (e271) - get paper here
  • Bauer, Aaron M.;Günther, Rainer 1995. An annotated type catalogue of the lacertids (Reptilia: Lacertidae) in the Zoological Museum, Berlin. Mitt. Zool. Mus. Berlin 71 (1): 37-62 - get paper here
  • Boulenger,G.A. 1920. Monograph of the Lacertidae, vol. 1. British Museum (London) - get paper here
  • Boycott, R.C. 1992. An Annotated Checklist of the Amphibians and Reptiles of Swaziland. The Conservation Trust of Swaziland - get paper here
  • Branch, William R. 1993. A Photographic Guide to Snakes and Other Reptiles of Southern Africa. Cape Town: Struik Publishers, 144 S.
  • Branch, William R. & Bauer, Aaron M. 2005. The herpetological contributions of Sir Andrew Smith. SSAR, 80 pp.
  • Broadley, D. G. & HOWELL, K. M. 1991. A check list of the reptiles of Tanzania, with synoptic keys. Syntarsus 1: 1—70
  • Broadley, D.G. 1962. On some reptile collections from the North-Western and North-Eastern Districts of Southern Rhodesia 1958-1961, with descriptions of four new lizards. Occ. Pap. Nat. Mus. South. Rhodesia 26 (B): 787-843
  • Broadley, D.G. 1972. A review of the Nucras tessellata group (Sauria: Lacertidae). Arnoldia 20: 1-35
  • Conradie, W. 2012. Geographic Distributions: Nucras taeniolata (Smith, 1838) Albany Sandveld Lizard. African Herp News (58): 21-22 - get paper here
  • CONRADIE, W., DOUCETTE-RIISE, S., VANHOOYDONCK, B., ENGELBRECHT, H., MEASEY, G.J., & TOLLEY, K. 2011. Herpetological survey of Rooipoort Nature Reserve, Northern Cape, South Africa. African Herp News (53): 35-41 - get paper here
  • CONRADIE, W., ENGELBRECHT, H., HERREL, A., MEASEY, G.J., NIELSEN, S.V., V ANHOOYDONCK, B., & TOLLEY, K.A. 2011. Reptile survey of Venetia Limpopo Nature Reserve, Limpopo Province - South Africa. African Herp News (55): 6-12 - get paper here
  • Conradie, Werner; Brian Reeves , Neil Brown, & Jan A. Venter 2016. Herpetofauna of the Oviston, Commando Drift and Tsolwana nature reserves in the arid interior of the Eastern Cape Province, South Africa. Indago 32: 81-98 - get paper here
  • Duméril, A. M. C. and G. Bibron. 1839. Erpétologie Générale on Histoire Naturelle Complète des Reptiles. Vol. 5. Roret/Fain et Thunot, Paris, 871 pp. - get paper here
  • Edwards, Shelley; Krystal A. Tolley, Bieke Vanhooydonck, G. John Measey and Anthony Herrel 2013. Is dietary niche breadth linked to morphology and performance in Sandveld lizards Nucras (Sauria: Lacertidae)? Biological Journal of the Linnean Society, DOI: 10.1111/bij.12148 - get paper here
  • FitzSimons, V.F. 1943. The lizards of South Africa. Transvaal Museum Memoir No.1 (Pretoria), 528 pp.
  • Gemel, R.; G. Gassner & S. Schweiger 2019. Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien – 2018. Ann. Naturhist. Mus. Wien, B 121: 33–248
  • Gray, J. E. 1864. Notes on some new lizards from south—eastern Africa, with the descriptions of several new species. Proc. Zool. Soc. London 1864: 58—62. - get paper here
  • Jacobsen, N. H. G. 1989. A herpetological survey of the Transvaal. Unpubl. thesis, Univ. Natal, South Africa
  • Pietersen, Darren, Verburgt, Luke & Davies, John 2021. Snakes and other reptiles of Zambia and Malawi. Struik Nature / Penguin Random House South Africa, 376 pp., ISBN 9781775847373
  • Smith, A. 1838. Contributions to the Natural History of Southern Africa. Art. VIII. Mag. Nat. Hist., London, 2 (14): 92-94
  • Spawls, S.; Howell, K.; Drewes, R.C. & Ashe, J. 2002. A field guide to the reptiles of East Africa. Academic Press, 543 pp. [reviews in HR 34: 396 and Afr. J. Herp. 51; 147] - get paper here
  • Steindachner,F. 1882. Über eine neue Eremias-Art aus dem Tale des Limpopo-(Krokodilflusses) in Transvaal. (E. holubi). Sitzungsb. Akad. Wien, 86: 83-85 - get paper here
  • Telford, N. S., ALEXANDER, G. J., BECKER, F. S., CONRADIE, W., JORDAAN, A., KEMP, L., ... & TOLLEY, K. A. 2022. EXTENSIONS TO THE KNOWN GEOGRAPHIC DISTRIBUTIONS OF REPTILES IN THE GREAT KAROO, SOUTH AFRICA. Herpetological Conservation and Biology, 17(1), 145-154 - get paper here
  • Tolley KA, Telford NS, Makhubo BG, Power RJ, Alexander GJ 2023. Filling the gap: Noteworthy herpetological discoveries in North West Province, South Africa. Zoosystematics and Evolution 99(1): 101-116 - get paper here
 
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