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Nucras boulengeri NEUMANN, 1900

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Higher TaxaLacertidae, Eremiadinae, Sauria, Lacertoidea, Squamata (lizards)
Subspecies 
Common NamesE: Uganda Savannah Lizard, Boulenger’s scrub lizard 
SynonymNucras boulengeri NEUMANN 1900: 56
Nucras emini BOULENGER 1907 (fide LOVERIDGE 1957: 230)
Nucras boulengeri — BOULENGER 1920: 16
Nucras emini — BOULENGER 1920: 10
Nucras kilosae LOVERIDGE 1922: 314 (fide BROADLEY & HOWELL 1991)
Nucras emini — LOVERIDGE 1923
Nucras boulengeri boulengeri — LOVERIDGE 1955: 176
Nucras boulengeri boulengeri — LOVERIDGE 1957: 7
Nucras boulengeri — BROADLEY & HOWELL 1991: 17
Nucras boulengeri kilosae — BISCHOFF 1991
Nucras boulengeri kilosae — BAUER & GÜNTHER 1995
Nucras boulengeri — HAAGNER et al. 2000
Nucras boulengeri — SPAWLS et al 2018: 192 
DistributionUganda, N Tanzania, Kenya, E/C Tanzania, Zambia

Type locality: Lubwa’s = Luba’s village, Busoga, Uganda.

Type locality: Tindiga, 5 miles south of Kilosa, Morogoro Dist., Tanzania [kilosae]  
Reproductionoviparous 
TypesType: BMNH 1946.8.6.16 (and possibly additional specimens).
Syntypes: BMNH [kilosae] 
DiagnosisDescription: Body elongate; head not depressed, its length (to ear opening) contained 4.5 to 5 times in the length from snout to vent; two postnasals; no granules between the supraoculars and the supraclliaries; interparietal not so long and narrow as in N. tessellata and N. Delalandii; occipital very small; subocular bordering the lip between the fourth and fifth upper labials; two supratemporals bordering the parietal; tympanum half as large as the ear-opening. Dorsal scales small, pointed behind, larger on the sides of the body; 45 to 53 scales round the body; ventrals in 6 longitudinal and 27 to 30 transverse series. Femoral pores 11 or 12. Foot much shorter than the head. Tail thinner than in N. tessedata and N. Delalandii, 1.25 to 1.33 as long as head and body.

Coloration: Colour brown above, with small indistinct blackish spots; bluish white beneath.

Size: This seems to be a much smaller species than the two previously known;

Distinctive characters: the rather large tympanum, the pointed dorsal scales, and the small foot. (Neumann 1900: 56)

Description of Nucras kilosae: Loveridge 1922: 314. 
CommentSynonymy: Broadley and Howell (1991) synonymized N. kilosae with N. boulengeri. 
EtymologyNamed after George Albert Boulenger (1858-1937), herpetologist at the British Museum of Natural History, London. For comments on the types of N. b. kilosae see BAUER & GÜNTHER 1995. 
References
  • Bauer, Aaron M.;Günther, Rainer 1995. An annotated type catalogue of the lacertids (Reptilia: Lacertidae) in the Zoological Museum, Berlin. Mitt. Zool. Mus. Berlin 71 (1): 37-62 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Boulenger, G.A. 1907. Descriptions of new lizards in the British Museum. Ann. Mag. Nat. Hist. (7) 19: 486-489. - get paper here
  • Boulenger,G.A. 1920. Monograph of the Lacertidae, vol. 1. British Museum (London) - get paper here
  • Broadley, D. G. & HOWELL, K. M. 1991. A check list of the reptiles of Tanzania, with synoptic keys. Syntarsus 1: 1—70
  • Edwards, Shelley; Krystal A. Tolley, Bieke Vanhooydonck, G. John Measey and Anthony Herrel 2013. Is dietary niche breadth linked to morphology and performance in Sandveld lizards Nucras (Sauria: Lacertidae)? Biological Journal of the Linnean Society, DOI: 10.1111/bij.12148 - get paper here
  • Haagner,G.V.; Branch,W.R. & Haagner,A.J.F. 2000. Notes on a collection of reptiles from Zambia and adjacent areas of the Democratic Republic of the Congo. Annals of the Eastern Cape Museum 1: 1 – 25
  • Loveridge, A. 1923. Notes on East African lizards collected 1920-1923 with the description of two new races of Agama lionotus Blgr. Proc. Zool. Soc. London 1923: 935-969 - get paper here
  • Loveridge, A. 1955. On a second collection of reptiles and amphibians taken in Tanganyika Territory by C. J. P. lonides,. Journ. E. Africa Nat. Hist. Soc. 22: 169-198
  • Loveridge, A. 1957. On a third collection of reptiles taken in Tanganyika by C.J.P. Ionides, Esq. Tanganyika Notes and Records 43: 1-19
  • Loveridge,A. 1922. New reptiles from Tanganyika Territory. Proc. Zool. Soc. London 1922: 313-315 - get paper here
  • Malonza, Patrick K.; David M. Mulwa, Joash O. Nyamache, Georgina Jones 2017. Biogeography of the Shimba Hills ecosystem herpetofauna in Kenya. Zoological Research 38(5): 1-11 - get paper here
  • Muchai, Vincent & Patrick Malonza 2011. The reptiles and amphibians of Kibwezi Forest, Ngulia and Chyulu Hills. Kenya Past and Present (39): 45-49 - get paper here
  • MÜLLER, J. 1998. Zur besseren Kenntnis von Nucras boulengeri (NEUMANN, 1900), einer wenig bekannten Eidechse aus Ostafrika. Die Eidechse 8 (3): 88-96 - get paper here
  • Neumann, O. 1900. Description of a new lizard of the genus Nucras from Usoga, British East Africa. Ann. Mag. Nat. Hist. (7) 5: 56 - get paper here
  • Pietersen, Darren, Verburgt, Luke & Davies, John 2021. Snakes and other reptiles of Zambia and Malawi. Struik Nature / Penguin Random House South Africa, 376 pp., ISBN 9781775847373
  • Razzetti, E. & Msuya, C.A. 2002. Field guide to the amphibians and reptiles of Arusha National Park (Tanzania). Publ. Ed. Negri Istituto, Oikos, Varese, 84 pp.
  • Spawls, S. & Rotich, D. 1997. An annotated checklist of the lizards of Kenya. J. East African Nat. Hist. 86: 61-83
  • Spawls, Steve; Kim Howell, Harald Hinkel, Michele Menegon 2018. Field Guide to East African Reptiles. Bloomsbury, 624 pp. - get paper here
 
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