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Oedura fimbria OLIVER & DOUGHTY, 2016

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Higher TaxaDiplodactylidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Western marbled velvet gecko 
SynonymOedura fimbria OLIVER & DOUGHTY 2016 
DistributionAustralia (Western Australia)

Type locality: Brockman Ridge, Western Australia, 23.3108°S, 119.9169°E, Western Australia  
TypesHolotype: WAM R154783, adult male with original tail, Brockman Ridge, Western Australia, 23.3108°S, 119.9169°E, Western Australia, collected by J. Fraser on 5 April 2004. Paratypes. All from Western Australia: WAM R105965, 7 km north of Mount Magnet (28.00°S, 117.88°E); WAM R119991, Hope Downs (23.01°S, 119.10°E); WAM R119993, Hope Downs (23.00°S, 119.12°E); WAM R129595, 120 km north-west of Newman (22.92°S, 118.88°E); WAM R129635, 120 km north-west of Newman (22.92°S, 119.02°E); WAM R135369, Mount Brockman (22.31°S, 117.32°E); WAM R154796–7, Walga Rock (27.40°S, 117.47°E); WAM R157504, WAM R157508, WAM R157516, Packsaddle Range (22.92°S, 118.89°E); WAM R157595, West Angelas (23.19°S, 118.86°E); WAM R160074, 32.5 km east-south-east of Meentheena Outcamp (21.33°S, 120.750°E); WAM R165150, 5 km north-north-west Python Pool (21.32°S, 117.23°E). 
DiagnosisDiagnosis. A large (mean SVL 94 mm, max 104 mm) species in the O. marmorata complex, with a moderately broad head (HW/SVL 0.18–0.21), moderately long body length (Trk/SVL 0.42–0.53), tail moderately long (TL/SVL 0.65–0.80), narrower than head and roughly circular in cross-section, rostral 25–60% divided, apical lamellae wide (TW/SVL 0.027–0.036), subdigital lamellae in a flared series wider than apical pair on fingers 3–4, 13–26 precloacal pores in adult males, and basic dorsal colouration (especially on juveniles) usually including 5 light transverse bands.

Comparisons. Oedura fimbria sp. nov. is most similar to O. cincta and shares moderately large size (SVL > 100 mm), moderately long tail which tends towards rounded in cross-section and is not wider than the head, wide flared subdigital lamellae, dorsal pattern usually consisting of numerous poorly-defined light flecks and blotches and often thin light bands. The only clear diagnostic morphological character between the two speices is an incomplete rostral crease on O. fimbria sp. nov. (25–60% versus 60 [rarely]–100% [usually] of the rostral height). There are some differences in average for relative head length and body length, but ranges overlap extensively (see Table 1).
Oedura fimbria sp. nov. can be distinguished from other lineages in the O. marmorata species complex from northern Australia by possessing a longer (TL/SVL 0.65–0.80 versus 0.53–0.63) and narrower tail (always narrower than the head). It differs from O. bella in its larger size (max SVL 104 mm versus 92 mm), longer tail (TL/SVL 0.65–0.80 versus 0.49–0.65), wider terminal lamellae (3TW/SVL 0.23–0.36 versus 0.21–0.30) and prominently flared subdigital lamellae series—especially on fingers 3 and 4 (proximal lamellae wider than apical lamellae versus not wider).
Oedura fimbria sp. nov. can be distinguished from the three other species of Oedura in the Kimberley region in Western Australia as follows: from O. gracilis by its moderately long and swollen tail (versus very long [approaching length of body] and tapering) and in having subdigital lamellae series that is flared around the midpoint of the digit (versus tapering); from O. filicipoda in having a narrow tail that is not wider than the head and near circular in cross-section (versus wider and very flattened) and smaller eyes (EYE/SVL 0.051–0.064 versus 0.071–0.076); and from O. murrumanu in having a smaller eye (EYE/SVL 0.051–0.064 versus 0.067–0.069) and a longer rostral crease (> 25% [typically 40–60%] versus < 25% of rostral height). 
CommentHabitat: relatively massive ranges, rocky outcrops and breakaways, caves and gorges, with a few occurrences on trees and one under tin. OLIVER & DOUGHTY 2016 have only observed this species on rocky outcrops with large boulders (Pilbara) or on cliff faces (Kennedy Range). 
EtymologyFimbria is Latin for fringe in reference to the lateral fringes of expanded lamellae along the sides of the digits. Used as a noun in apposition. 
  • Ellis, Ryan J.; Paul Doughty and Aaron M. Bauer 2018. An annotated type catalogue of the geckos and pygopods (Squamata: Gekkota: Carphodactylidae, Diplodactylidae, Gekkonidae, Pygopodidae) in the collection of the Western Australian Museum. Records of the Western Australian Museum 33: 051–094 - get paper here
  • OLIVER, PAUL M.; PAUL DOUGHTY 2016. Systematic revision of the marbled velvet geckos (Oedura marmorata species complex, Diplodactylidae) from the Australian arid and semi-arid zones. Zootaxa 4088 (2): 151–176 - get paper here
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