Oligodon macrurus (ANGEL, 1927)

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Higher Taxa	Colubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies
Common Names	E: Angel's Kukri Snake
Synonym	Simotes violaceus macrurus ANGEL 1927 Holarchus violaceus macrurus – BOURRET 1936 Oligodon macrurus — SMITH 1935: 207 Oligodon macrurus— SMITH 1943: 207 Oligodon macrurus — NGUYEN et al. 2009 Oligodon macrurus — WALLACH et al. 2014: 499 Oligodon arenarius VASSILIEVA 2015 Oligodon macrurus — YUSHCHENKO et al. 2023
Distribution	Vietnam (Ninh Thuan, Binh Thuan) Type locality: Pointe Lagan, Southern Annam (= Ninh Thuan Province, Vietnam) Neotype locality: “coastal dune area about 1 km from the coast line, Mui Ne, Binh Thuan Province, Vietnam (near 10°57’59.6″N, 108°19’46.5″E)” [DOR] arenarius: S Vietnam (Ba Ria–Vung Tau); Type locality: Binh Chau–Phuoc Buu Nature Reserve, Xuen Moc District, Ba Ria–Vung Tau Province, southern Vietnam, coordinates 10°29'46"N, 107°27'54"E, elevation 5 m elevation.
Reproduction	oviparous
Types	Neotype: ZFMK 88885, collected by Peter Geissler, April 27, 2009 (designated by Yushchenko et al. 2023) Holotype: lost, was MNHN-RA 1928.0119 (but no longer found, Sang et al. 2009). Not located in MNHN-RA collections and thus considered lost (pers. comm., I. Ineich, March 2019, 30 Nov 2021). Holotype: ZMMU R-14503 (field ID ZMMU ABV–00813), adult male, collected by Anna B. Vassilieva on 14 November 2014. Paratypes. ZMMU R-14002 (field ID ZMMU NAP-03137), adult male from type locality: Binh Chau–Phuoc Buu Nature Reserve, Xuen Moc Disctrict, Ba Ria–Vung Tau Province, southern Vietnam (10°32'18"N, 107°28'37"E, 43 m a.s.l.), collected on 8 November 2010 by Anna B. Vassilieva; ZMMU R-14002 (field ID ZMMU NAP-03138), adult female from type locality (10°29'50"N, 107°27'58"E, 5 m a.s.l.) collected on 9 November 2010 by Anna B. Vassilieva; VNMN 04724 (field ID NAP-03884), adult female from type locality, environs of the resort Ho Coc (10°29'54"N, 107°28'08"E; 8 m a.s.l.), captured on 10 July 2012 by local people and collected by Anna B. Vassilieva and Nikolay Poyarkov; ZMMU R-14504, adult female from type locality (10°29'41"N, 107°27'46"E; 7 m a.s.l.), collected on 16 November 2014 by Anna B. Vassilieva [arenarius]
Diagnosis	Additional details, e.g. a detailed description or comparisons (19489 characters), are available for collaborators and contributors. Please contact us for details.
Comment	Synonymy: YUSHCHENKO et al. 2023 synonymized Oligodon arenarius with Oligodon macrurus. Etymology: O. arenarius was named after the Latin word "arena" meaning "sand", "sandy land", nominative case, masculine gender. The name is intended to reflect the evident preference by the new species for sandy coastal habitats and the snake's remarkable ability to plunge into the sand and move under it.
References	Angel, F. 1927. Liste des reptiles et des batracien rapportés d’Indo-Chine par M.P. Chevey. Description d’une variété nouvelle de Simotes violaceus Cantor. Bull. Mus. natl. Hist. nat. Paris 33: 496-498 - get paper here DAVID, PATRICK; TRUONG QUANG NGUYEN, TAO THIEN NGUYEN, KE JIANG, TIANBO CHEN, ALEXANDRE TEYNIÉ & THOMAS ZIEGLER 2012. A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from northern Vietnam, southern China and central Laos. Zootaxa 3498: 45–62 - get paper here Geissler, Peter; Alexander R. Krohn, Daniel Rennert 2011. Herpetofaunal records in coastal dune areas, Binh Thuan Province, Southern Vietnam, with the rediscovery of Oligodon macrurus Angel, 1927. Russ. J. Herpetol. 18 (4): 317–324 - get paper here NEANG, THY; L. LEE GRISMER & JENNIFER C. DALTRY 2012. A new species of kukri snake (Colubridae: Oligodon Fitzinger, 1826) from the Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa 3388: 41–55 - get paper here Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp. Nguyen, Sang Ngoc; Vu Dang Hoang Nguyen, Robert W. Murphy 2021. Notes on the hemipenis and head scalation of Oligodon arenarius Vassilieva, 2015 (Serpentes, Colubridae). Herpetology Notes 14: 91-94 NGUYEN, SANG NGOC; VU DANG HOANG NGUYEN, SON HONG LE, ROBERT W. MURPHY 2016. A new species of kukri snake (Squamata: Colubridae: Oligodon Fitzinger, 1826) from Con Dao Islands, southern Vietnam. Zootaxa 4139 (2): 261–273 - get paper here Smith, M.A. 1943. The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. 3 (Serpentes). Taylor and Francis, London. 583 pp. VASSILIEVA, ANNA B. 2015. A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from coastal southern Vietnam. Zootaxa 4058 (2): 211–226 - get paper here VASSILIEVA, ANNA B.; PETER GEISSLER, EDUARD A. GALOYAN, NIKOLAY Jr A. POYARKOV, ROBERT WAYNE VAN DEVENDER, WOLFGANG BÖHME 2013. A new species of Kukri Snake (Oligodon Fitzinger, 1826; Squamata: Colubridae) from the Cat Tien National Park, southern Vietnam. Zootaxa 3702 (3): 233–246 - get paper here Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp. Yushchenko PV, Lee JL, Pham HM, Geissler P, Syromyatnikova EV, Poyarkov Jr. NA 2023. The taxonomic status of the kukri snake Oligodon arenarius Vassilieva, 2015 with a redescription of Oligodon macrurus (Angel, 1927) (Squamata, Serpentes, Colubridae). Vertebrate Zoology 73: 97-125 - get paper here
External links	IUCN Red List National Center for Biotechnology Information https://www.facebook.com/groups/stshc/posts/557850499772886/ Google images

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