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Oligodon nagao DAVID, NGUYEN, NGUYEN, JIANG, CHEN, TEYNIÉ & ZIEGLER, 2012

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesChinese: 方斑小头蛇 (Fang Ban Xiao Tou She)
E: Nagao Kukri Snake
Vietnamese: Rắn khiếm na-gao 
SynonymOligodon nagao DAVID, NGUYEN, NGUYEN, JIANG, CHEN, TEYNIÉ & ZIEGLER 2012
Oligodon nagao — WALLACH et al. 2014: 500
Oligodon nagao — DAVID et al. 2022 
DistributionN Vietnam (Cao Bang, Ninh Binh), Laos (Khammouane), SW China (Guangxi)

Type locality: Huu Lien forest, Huu Lung District, Lang Son Province, Vietnam, at elevation of about 300 m elevation.  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: VNMN A.2012.1, adult male; collected by Tao Thien Nguyen on 30 June 2009. Paratypes. Four specimens, all adult males: MNHN-RA 2012.0216, from Huu Lien forest, Huu Lung District, Lang Son Province, Vietnam, at elevation of about 300 m a.s.l.; collected by Tao Thien Nguyen, 2 July 2009.—IEBR A.2012.6, from Duc Quang Commune, Ha Lang District, Cao Bang Province, Vietnam; collected by Truong Quang Nguyen et al., 14 October 2011 (22o42.859’N, 106o39.853’E, elevation 476 m).—KIZ 014591, from Nonggang National Nature Reserve, Longzhou County, Guangxi Autonomous Region, People’s Republic of China; collected by Tianbo Chen, October, 2011.—ZFMK 93281, from Ban Nathan, Hin Boun District, Khammouane Province, Laos PDR (17o58.854’N, 104o49.517’E, elevation 172 m); collected by Alexandre Teynié et al., 17 May 2012, during the mission “Opération Canopée, Inventaire de la Biodiversité Forestière du Laos 2012-2015”. 
DiagnosisDiagnosis: A large species (TL up to at least 786 mm) of the genus Oligodon characterized by the combination of (1) 9 or 10 maxillary teeth, the last three or four strongly enlarged, (2) hemipenes not forked but divided into two lobes, thick and bulbous, reaching in situ the 16th SC, smooth, each lobe with a papilla, (3) 17–17–15 (or 17–15–15 DSR in one specimen); (4) cloacal plate entire, (5) complete complement of head scales, including 1 loreal on each side, (6) 8 supralabials (7 in one specimen), fourth and fifth (third and fourth in one specimen) entering orbit, (7) 184–193 ventrals in five males (females unknown), (8) tail relatively short in males (ratio TaL/TL: 0.135–0.146), (9) dorsal pattern made of a dark background colour with 27–37 darker, pale centered, butterfly-shaped blotches on the body and 5–8 on the tail, and (10) venter cream, heavily marked with dark pigmentation. 
Comment 
EtymologyNamed after the Nagao Natural Environment Foundation, Japan, for the support of natural sciences research and conservation in developing countries in Asia. 
References
  • Böhme, Wolfgang 2014. Herpetology in Bonn. Mertensiella 21. vi + 256 pp. - get paper here
  • David, Patrick; Sjon Hauser & Gernot Vogel 2022. A NEW SPECIES OF THE GENUS Oligodon FITZINGER, 1826 (REPTILIA: COLUBRIDAE) FROM SOUTHERN LAOS TAPROBANICA 11 (1): 12-24 - get paper here
  • DAVID, PATRICK; TRUONG QUANG NGUYEN, TAO THIEN NGUYEN, KE JIANG, TIANBO CHEN, ALEXANDRE TEYNIÉ & THOMAS ZIEGLER 2012. A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from northern Vietnam, southern China and central Laos. Zootaxa 3498: 45–62 - get paper here
  • NGUYEN, QUANG HUY et al. 2022. Geographic distribution: OLIGODON NAGAO (Nagao Kuri Snake). VIETNAM: NINH BINH PROVINCE. Herpetological Review 53 (1): 80
  • VASSILIEVA, ANNA B.; PETER GEISSLER, EDUARD A. GALOYAN, NIKOLAY Jr A. POYARKOV, ROBERT WAYNE VAN DEVENDER, WOLFGANG BÖHME 2013. A new species of Kukri Snake (Oligodon Fitzinger, 1826; Squamata: Colubridae) from the Cat Tien National Park, southern Vietnam. Zootaxa 3702 (3): 233–246 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Wang, Kai; Jinlong Ren, Hongman Chen, Zhitong Lyu, Xianguang Guo Ke Jiang, Jinmin Chen, Jiatang Li, Peng Guo, Yingyong Wang, Jing Che 2020. The updated checklists of amphibians and reptiles of China. Biodiversity Science 28 (2): 189-218 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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