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Oreosaurus achlyens (UZZELL, 1958)

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Higher TaxaGymnophthalmidae (Cercosaurinae), Sauria, Gymnophthalmoidea, Squamata (lizards)
Subspecies 
Common Names 
SynonymProctoporus achlyens UZZELL 1958
Proctoporus achlyens — PETERS & DONOSO-BARROS 1970: 238
Proctoporus achlyens — DOAN & SCHARGEL 2003
Riama achlyens — DOAN & CASTOE 2005
Oreosaurus achlyens — SÁNCHEZ-PACHECO et al. 2017 
DistributionVenezuela (Aragua), elevation 1100-1150

Type locality: Maracay-Rancho Grande Road, Estado Aragua, Venezuela, elevation 1150 m.  
Reproductionoviparous 
TypesHolotype: UMMZ 117333 
Diagnosis 
CommentSynonymy: note that Oreosaurus is different from Oreosaura Barabanov & Ananjeva 2007, a subgenus of Phrynocephalus. 
EtymologyNamed after Greek achlyoo (ἀχλυόω), darken, make dim; become gloomy, grow dark. [“...The name achlyens is from the Greek, ἀχλυόω, ‘to grow dark.’...”]. (from Esteban Lavilla, pers. comm., May 2024) 
References
  • Doan, T. M. & Castoe, T.A. 2005. Phylogenetic taxonomy of the Cercosaurini (Squamata: Gymnophthalmidae), with new genera for species of Neusticurus and Proctoporus. Zoological Journal of the Linnean Society, 143: 405–416. - get paper here
  • Doan, T.M. 2003. A south-to-north biogeographic hypothesis for Andean speciation: evidence from the lizard genus Proctoporus (Reptilia, Gymnophthalmidae). Journal of Biogeography 30: 361–374 - get paper here
  • Doan, T.M. & Schargel, W.E. 2003. Bridging the gap in Proctoporus distribution: a new species (Squamata: Gymnophthalmidae) from the Andes of Venezuela. Herpetologica 59 (1): 68-75 - get paper here
  • Peters, James A. & Donoso-Barros, Roberto 1970. Catalogue of the Neotropical Squamata: Part II. Lizards and Amphisbaenians. Bull. US Natl. Mus. 297: 293 pp. - get paper here
  • RIVAS, G. A., NUNES, P. M. S., BARAN, A., JOWERS, M. J., SMITH, E. N., HERNÁNDEZ-MORALES, C., & SCHARGEL, W. E. 2021. A new species of Oreosaurus (Squamata: Gymnophthalmidae) from the Turimiquire Massif, northeastern Venezuela. Zootaxa 5023 (4): 571-584 - get paper here
  • Rivas, G.; Schargel, W.E. & Meik, J.M. 2005. A new species of Riama (Squamata: Gymnophthalmidae), endemic to the Península de Paria, Venezuela. Herpetologica 61 (4): 461-468 - get paper here
  • RIVAS, GILSON A.; CÉSAR R. MOLINA, GABRIEL N. UGUETO, TITO R. BARROS, CÉSAR L. BAR- RIO-AMORÓS & PHILIPPE J. R. KOK 2012. Reptiles of Venezuela: an updated and commented checklist. Zootaxa 3211: 1–64 - get paper here
  • Roth, W.D. & Gans, C. 1960. The luminous organs of Proctoporus (Sauria, Reptilia) - a a re-evaluation. Breviora (125): 1-12 - get paper here
  • Sánchez-Pacheco, S. J., Torres-Carvajal, O., Aguirre-Peñafiel, V., Sales-Nunes, P. M., Verrastro, L., Rivas, G. A., Rodrigues, M. T., Grant, T. and Murphy, R. W. 2017. Phylogeny of Riama (Squamata: Gymnophthalmidae), impact of phenotypic evidence on molecular datasets, and the origin of the Sierra Nevada de Santa Marta endemic fauna. Cladistics, doi:10.1111/cla.12203 [print: 2018] - get paper here
  • Sánchez-Pacheco, Santiago J.; David A. Kizirian, and Pedro M. Sales-Nunes 2011. A New Species of Riama from Ecuador Previously Referred to as Riama hyposticta (Boulenger, 1902) (Squamata: Gymnophthalmidae). American Museum Novitates (3719): 1-15 - get paper here
  • Test, Frederic H.;Sexton, Owen J.;Heatwole, Harold 1966. Reptiles of Rancho Grande and vicinity, Estado Aragua, Venezuela. Miscellaneous publications, Museum of Zoology, University of Michigan (128): 1-63 - get paper here
  • Uzzell, T.M. 1958. Teiid lizards related to Proctoporus luctuosus, with the description of a new species from Venezuela. Occasional Papers of the Museum of Zoology University of Michigan 597: 1-15 - get paper here
 
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