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Pachydactylus werneri HEWITT, 1935

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Werner's Thick-toed Gecko 
SynonymPachydactylus capensis werneri HEWITT 1935: 315
Pachydactylus capensis weberi HEWITT 1935: 315
[Pachydactylus capensis gariesensis — HEWITT 1935: 315
Pachydactylus werneri — FITZSIMONS 1943: 85
Pachydactylus weberi werneri — LOVERIDGE 1947: 394
Pachydactylus werneri — LOVERIDGE 1947: 394
Pachydactylus werneri — WERMUTH 1965: 124
Pachydactylus werneri — BAUER et al. 2006
Pachydactylus werneri — CONRADIE et al. 2019 
DistributionNamibia (Maltahöhe, Swakopmund, Karibib Districts); elevation usually below 1000 m.

Type locality: Khan River, north of Walfisch Bay, Southwest Africa.  
TypesLectotype: PEM R 16049 (formerly AM 6613 (part)), coll. R. D. Bradfield (designated by BAUER et al. 2006). 
DiagnosisDIAGNOSIS.—To 52.6 mm SVL. Pachydactylus werneri may be distinguished from all other members of the P. serval/weberi group by the combination of the following characters: rostral (and in some cases first supralabial) excluded from nostril; supranasals in variable contact; nostril rims distinctly raised; eyes very large, rostrum pointed; scales on dorsum of head granular, those on snout much larger than those of interorbital region; dorsal scalation heterogewith relatively small keeled tubercles arranged in 16–18 regular rows; tubercle tips and/or keels often white; thighs with few, small, scattered, keeled to mucronate tubercles; limbs and toes very elongate, slender, toe pads wide; typically 5 undivided lamellae beneath digit IVof pes; tail to approximately 90% SVL; moderately annulate, bearing whorls of small, keeled, strongly pointed, white-tipped tubercles; adult pattern of diffuse, dark-edged light bands: one on nape, one behind axilla, one on mid-trunk, and one anterior to sacrum; pale bands often not, or barely, lighter than grayish- or pinkish-brown background coloration, pattern often disrupted and appearing as a series of irregular brown cross bands or spots; juvenile pattern similar to adult with four light cross bands (Girard 2002), although often only the nape band is prominent [from BAUER et al. 2006].
CommentSynonymy partly after WERMUTH 1965 and BAUER et al. (2006).

Specimens from other areas in Namibia have been described as new species by BAUER et al. (2006).

Distribution: see map in BRANCH et al. 2011.

HABITAT: The specimens of Pachydactylus werneri that BAUER et al. (2006) collected were active at night on rock faces along the north bank of the Swakop River, and Mertens (1955) found a specimen under a stone. However, according to Mirko Barts (pers. comm., May 2004 cited in BAUER et al. 2006) the species is not typically rupicolous and is chiefly active on riverine vegetation. 
  • Bauer, Aaron M. Lamb, Trip. Branch, William R. 2006. A revision of the Pachydactylus serval and P. weberi groups (Reptilia: Gekkota: Gekkonidae) of Southern Africa, with the description of eight new species. Proc. Cal. Acad. Sci. 57 (12-24):595-709. - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Branch, William R.; Aaron M. Bauer, Todd R. Jackman, and Matthew Heinicke 2011. A New Species of the Pachydactylus weberi Complex (Reptilia: Squamata: Gekkonidae) from the Namibrand Reserve, Southern Namibia. Breviora (524): 1-15. - get paper here
  • CONRADIE, WERNER; WILLIAM R. BRANCH, & GILLIAN WATSON 2019. Type specimens in the Port Elizabeth Museum, South Africa, including the historically important Albany Museum collection. Part 2: Reptiles (Squamata). Zootaxa 4576 (1): 001–045 - get paper here
  • Girard, Francis 2002. Maintenance and Captive Reproduction of Two Small, South African Geckos Pachydactylus geitje (SPARRMAN, 1778) and Pachydactylus weberi Raux, 1907. Gekko 3 (1): 14-18
  • Herrmann, H.-W.; W.R. Branch 2013. Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist. Journal of Arid Environments 93: 94–115 - get paper here
  • Hewitt, J. 1935. Some new forms of batrachians and reptiles from South Africa. Rec. Albany Mus. 4: 283-357
  • Loveridge, A. 1947. Revision of the African lizards of the family Gekkondiae. Bull. Mus. Comp. Zool. Harvard 98: 1-469 - get paper here
  • Wermuth, H. 1965. Liste der rezenten Amphibien und Reptilien. Gekkonidae, Pygopodidae, Xantusiidae. Das Tierreich (80): 1—246
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