Panolopus lanceolatus SCHOOLS & HEDGES, 2024
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Panolopus lanceolatus
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| Higher Taxa | Diploglossidae, Diploglossa, Anguimorpha, Sauria, Squamata (lizards) |
| Subspecies | |
| Common Names | E: westcentral Smooth-scaled Forest Lizard |
| Synonym | Panolopus lanceolatus SCHOOLS & HEDGES 2024: 170 Celestus costatus — SCHWARTZ & HENDERSON, 1991:368 (part) Celestus costatus — POWELL et al., 1999:104 (part) Celestus costatus — HEDGES et al., 2019:16 (part) Panolopus costatus — SCHOOLS & HEDGES, 2021:230 (part) Panolopus costatus — LANDESTOY et al., 2022:205 (part) |
| Distribution | Haiti Type locality: intersection between Ennery and Plaisance, a couple hundred meters E on intersecting road (19.52387, -72.45154; 925 m). |
| Reproduction | |
| Types | Holotype. MCZ R-190692, an adult female from first and only major intersection between Ennery and Plaisance, a couple hundred meters E on intersecting road, collected by D. Luke Mahler and Richard E. Glor on 6 August 2009 (19.52387, -72.45154; 925 m). Paratypes (n=8). HAITI. Artibonite. KU 226436–9, 1.2 mi w Ennery, 1977. Nord. MCZ R-63383–4, Austin Stanley Rand and James Draper Lazell, Dondon, 16 July 1960; MCZ R-63386, Austin Stanley Rand, James Draper Lazell, Jr., Jean Bernard between Cap Haitien and Grande Riviere du Nord, 16 July 1960; MCZ R-190691, D. Luke Mahler, Richard E. Glor, first and only major intersection between Ennery and Plaisance, couple hundred meters E on intersecting road, 6 August 2009. |
| Diagnosis | Diagnosis: Panolopus lanceolatus sp. nov. has (1) a dorsal pattern of absent/irregular dots/dots in series/dots in chevrons, (2) head markings absent/present, (3) markings in the longitudinal paramedian area present, (4) dots arranged in bars in the lateral band present, (5) an adult SVL of 78.5–104 mm, (6) ventral scale rows, 93–102, (7) midbody scale rows, 37–43, (8) total lamellae on one hand, 41–52, (9) total strigae on ten scales, 186–234, (10) relative length of all digits on one hindlimb, 28.4–35.9 %, (11) relative distance between the angled subocular and mouth, 0.567–0.704 %, (12) relative eye length, 3.01–3.51 %, (13) relative forelimb length, 17.4–23.9 %, (14) relative ear width, 0.770–1.35 %, (15) relative rostral height, 1.78–2.28 %, (16) relative head length, 15.2–18.2 %, (17) relative mental width, 1.45–1.96 %, (18) relative postmental width, 2.36–2.66 %, (19) relative cloacal width, 8.01–8.76 %, (20) relative prefrontal width, 3.97–4.55 %, (21) relative largest supraocular width, 2.20–2.71 %, (22) relative longest finger length, 4.76–6.36 %, (23) relative distance between the ear and eye, 6.45–7.70 %, (24) relative head width, 68.0–77.6 %, (25) relative frontal width, 63.1–72.1 %, (26) relative nasal height, 0.904–1.06 %, (27) relative angled subocular height, 0.484–0.854 %, (28) relative distance between the eye and naris, 4.58–5.05 %, (29) relative canthal iii length, 1.73–2.00 %, (30) relative angled subocular width, 2.38–3.09 %, and (31) relative nasal length, 1.48–1.95 %. No genetic data are available for estimating the species stem time or crown time. we distinguish Panolopus lanceolatus sp. nov. from the other species of Panolopus based on a complex of traits. From Panolopus aenetergum, we distinguish P. lanceolatus sp. nov. by the longitudinal paramedian lines (present versus absent), the ventral scale rows (93–102 versus 80–86), the midbody scale rows (37–43 versus 35–36), the total lamellae on one hand (41–52 versus 40), the total strigae on ten scales (186–234 versus 267), the relative length of digits on one hindlimb (28.4–35.9 versus 26.8), the relative distance between angled subocular and mouth (0.567– 0.704 versus 0.717), the relative cloacal width (8.01–8.76 versus 7.60), the relative distance between the ear and eye (6.45–7.70 versus 8.40), the relative frontal width (63.1–72.1 versus 88.2), the relative nasal height (0.904–1.06 versus 1.15), the relative angled subocular height (0.484–0.854 versus 1.08), the relative distance between the eye and naris (4.58–5.05 versus 4.35), and the relative angled subocular width (2.38–3.09 versus 2.07). From P. aporus, we distinguish P. lanceolatus sp. nov. by the relative largest supraocular width (2.20–2.71 versus 2.74–3.62) and the relative nasal height (0.904–1.06 versus 1.08–1.23). From P. chalcorhabdus, we distinguish P. lanceolatus sp. nov. by the angled subocular height (3.29–5.91 versus 2.77–3.28) (see Remarks). From P. costatus, we distinguish P. lanceolatus sp. nov. by the relative distance between the eye and naris (4.58–5.05 versus 5.08–5.50). From P. curtissi, we distinguish P. lanceolatus sp. nov. by the dots arranged in bars in the lateral areas (present versus absent), the total lamellae on one hand (41–52 versus 32–39), the relative length of digits on one hindlimb (28.4–35.9 versus 20.8–28.1), and the relative longest finger length (4.76–6.36 versus 3.59–4.54). From P. diastatus, we distinguish P. lanceolatus sp. nov. by the longitudinal paramedian lines (present versus absent) and the relative length of digits on one hindlimb (28.4–35.9 versus 21.5–27.4). From P. emys, we distinguish P. lanceolatus sp. nov. by the total strigae on ten scales (186–234 versus 238–311) and the relative angled subocular width (2.38–3.09 versus 2.12–2.20). From P. hylonomus, we distinguish P. lanceolatus sp. nov. by the dots arranged in bars in the lateral areas (present versus absent), the adult SVL (78.5–104 versus 59.3–76.5), the relative length of digits on one hindlimb (28.4–35.9 versus 22.8–28.2), and the relative postmental width (2.36–2.66 versus 2.67–2.89). From P. lapierrae sp. nov., we distinguish P. lanceolatus sp. nov. by the relative postmental width (2.36–2.66 versus 2.68–3.58), the relative prefrontal width (3.97–4.55 versus 4.73–4.75), the relative longest finger length (4.76–6.36 versus 4.49–4.55), the relative distance between the ear and eye (6.45–7.70 versus 7.78–8.43), the relative head width (68.0–77.6 versus 77.7–78.1), the relative frontal width (63.1–72.1 versus 77.6–79.0), and the relative distance between the eye and naris (4.58–5.05 versus 5.21). From P. leionotus, we distinguish P. lanceolatus sp. nov. by the frontal width by the SVL (3.48–3.84 versus 3.89–5.00) (see Remarks). From P. marcanoi, we distinguish P. lanceolatus sp. nov. by the relative nasal height (0.904–1.06 versus 1.07–1.28). From P. melanchrous, we distinguish P. lanceolatus sp. nov. by the relative postmental width (2.36–2.66 versus 2.71–3.38). From P. neiba, we distinguish P. lanceolatus sp. nov. by the frontal width by the SVL (3.48–3.84 versus 3.86–4.52) (see Remarks). From P. nesobous, we distinguish P. lanceolatus sp. nov. by the relative ear width (0.770–1.35 versus 1.39–1.60), the relative postmental width (2.36– 2.66 versus 2.82–3.02), the relative prefrontal width (3.97–4.55 versus 4.74–4.81), the relative largest supraocular width (2.20–2.71 versus 2.85–3.11), the relative distance between the ear and eye (6.45–7.70 versus 7.91–10.0), the relative nasal height (0.904–1.06 versus 1.12–1.14), the relative distance between the eye and naris (4.58– 5.05 versus 5.62–5.73), and the relative width of canthal iii (1.73–2.00 versus 2.01–2.12). From P. oreistes, we distinguish P. lanceolatus sp. nov. by the relative distance between the eye and naris (4.58–4.98 versus 5.01–5.63). From P. psychonothes, we distinguish P. lanceolatus sp. nov. by the relative nasal height (0.904–1.06 versus 1.12– 1.32). From P. saonae, we distinguish P. lanceolatus sp. nov. by the longitudinal paramedian lines (present versus absent), the relative largest supraocular width (2.20–2.71 versus 2.77), the relative frontal width (63.1–72.1 versus 72.5), the relative distance between the eye and naris (4.58–5.05 versus 6.43), and the relative angled subocular width (2.38–3.09 versus 2.31). From P. semitaeniatus sp. nov., we distinguish P. lanceolatus sp. nov. the relative ear width (0.770–1.35 versus 1.90–2.30), the relative rostral height (1.78–2.28 versus 2.41–2.63), the relative head width (68.0–77.6 versus 58.8–63.8), and the relative nasal height (0.904–1.06 versus 1.08). From P. unicolor sp. nov., we distinguish P. lanceolatus sp. nov. by the adult SVL (78.5–104 versus 67.6), the total strigae on ten scales (186–234 versus 144), the relative length of digits on one hindlimb (28.4–35.9 versus 36.8), the relative distance between angled subocular and mouth (0.567–0.704 versus 0.533), the relative ear width (0.770–1.35 versus 1.60), the relative postmental width (2.36–2.66 versus 2.80), the relative cloacal width (8.01–8.76 versus 7.61), the relative prefrontal width (3.97–4.55 versus 4.69), the relative largest supraocular width (2.20–2.71 versus 3.12), the relative longest finger length (4.76–6.36 versus 6.65), the relative frontal width (63.1–72.1 versus 58.2), the relative nasal height (0.904–1.06 versus 1.15), the relative distance between the eye and naris (4.58–5.05 versus 5.52), and the relative nasal width (1.48–1.95 versus 2.00). (Schools & Hedges 2024) Additional details (753 characters) are available for collaborators and contributors. 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| Comment | Distribution: for a map see Schools & Hedges 2024: 134 (Fig. 50) Similar species: Panolopus lanceolatus and P. chalcorhabdus cannot be morphologically separated based on standard characters; however, they can be separated by the angled subocular width divided by the angled subocular height (3.29–5.91 [n=7] versus 2.77–3.28 [n=5]). Panolopus lanceolatus sp. nov. and P. leionotus cannot be morphologically separated based on standard characters; however, they can be separated by the frontal width divided by the SVL (3.48–3.84 [n=8] versus 3.89–5.00 [n=5]). Panolopus lanceolatus sp. nov. and P. neiba cannot be morphologically separated based on standard characters; however, they can be separated by the frontal width divided by the SVL (3.48–3.84 [n=8] versus 3.86–4.52 [n=5]). (Schools & Hedges 2024) Synonymy: from SCHOOLS & HEDGES 2024. |
| Etymology | Named after Latin lanceolatus, a feminine adjective, meaning spear-like, referring to the flat and pointed head of this species. |
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