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Pantherophis quadrivittatus (HOLBROOK, 1836)

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Higher TaxaColubridae, Colubrinae, Lampropeltini, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Yellow Rat Snake (quadrivittata) 
SynonymColuber reticularis DAUDIN 1803 (part., see comment)
Coluber quadrivittatus HOLBROOK 1836
Coluber quadri-vittatus HOLBROOK 1842 (fide DUMÉRIL et al. 1854)
Scotophis quadrivittatus — BAIRD & GIRARD 1853: 80
Elaphis quadri-vittatus — DUMÉRIL, BIBRON & DUMÉRIL 1854: 265
Elaphis holbrookii DUMÉRIL & BIBRON 1854: 272
Elaphis quadrivittatus — GARMAN 1884: 56
Coluber obsoletus lemniscatus COPE 1888: 386 (part.)
Callopeltis quadrivittatus — LOENNBERG 1894
Coluber quadrivittatus— COPE 1900: 838
Elaphe quadrivittata deckerti BRADY 1932
Elaphe williamsi BARBOUR & CARR 1940: 340 (part.)
Elaphe quadrivittata parallela BARBOUR & ENGELS 1942: 103
Elaphe quadrivittata — LEWIS 1946
Elaphe obsoleta rossalleni NEILL 1949
Elaphe obsoleta deckerti — STANKOWSKI 1986
Elaphe obsoleta obsoleta — CONANT & COLLINS 1991: 197
Elaphe obsoleta quadrivittata — CONANT & COLLINS 1991: 198
Elaphe obsoleta rossalleni — CONANT & COLLINS 1991: 198
Elaphe obsoleta obsoleta — CROTHER 2000: 62
Elaphe obsoleta quadrivittata — CROTHER 2000: 62
Elaphe obsoleta rossalleni — CROTHER 2000: 62
Elaphe obsoleta rossalleni — TENNANT & BARTLETT 2000: 385 
DistributionUSA (Virginia, South Carolina, North Carolina, Georgia [east of the Chattohoochee River], Florida [east of the Apalachicola River]).  
Reproductionoviparous 
TypesHolotype: ANSP 3773; South Carolina (?) [quadrivittatus] 
DiagnosisAdditional details, e.g. a detailed description or comparisons (624 characters), are available for collaborators and contributors. Please contact us for details. 
CommentDistribution: see map in Burbrink et al. 2021: 539 (Fig.1).

The diagnosis of E. alleghaniensis by BURBRINK (2001) is unfortunately not very useful as it gives mostly relative characters (”E. alleghaniensis tends to have a higher number of subcaudals than E. spiloides” etc.) and hardly any comparisons with other species.

Phylogenetics: Burbrink et al. 2020 recognized three species (P. obsoletus (western clade), P. alleghaniensis (eastern), and P. spiloides (central ) but then discovered that they type of P. alleghaniensis actually was within the central clade (only noted in the supplemental material). The correct clade names are thus: P. obsoletus (western; west of the Mississippi River), P. alleghaniensis (central), and P. quadrivittatus (eastern) (T. Taggart, pers. comm., 20 Oct 2021, following Burbrink et al. 2021).

Habitat: partly arboreal (Harrington et al. 2018). 
EtymologyNamed after the Allegheny Mountains (a part of the Apalachians) or the Allegheny River in west-central Pennsylvania. The meaning of the word, which comes from the Lenape (Delaware) Native Americans, is not definitively known but is usually translated as "fine river" (Wikipedia). 
References
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