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Pareas berdmorei THEOBALD, 1868

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Higher TaxaPareidae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
SubspeciesPareas berdmorei berdmorei THEOBALD 1868
Pareas berdmorei truongsonicus POYARKOV et al. 2022
Pareas berdmorei unicolor (BOURRET 1934) 
Common NamesE: Mengla Snail-eating Snake
Chinese: 勐腊钝头蛇 (Mengla Dun-tou-she) 
SynonymPareas berdmorei THEOBALD 1868
Pareas menglaensis WANG, CHE, LIU, KI, JIN, JIANG, SHI & GUO 2020
Pareas berdmorei — DAVID & DEUTI 2022
Pareas menglaensis — WANG et al. 2022
Pareas (Pareas) berdmorei — POYARKOV et al. 2022: 45

Pareas berdmorei unicolor (BOURRET 1934)
Amblycephalus carinatus unicolor BOURRET 1934
Pareas carinatus unicolor — NGUYEN et al. 2009
Pareas (Pareas) berdmorei unicolor — POYARKOV et al. 2022 
DistributionMyanmar, China, Vietnam, Laos, Thailand

Type locality: Mon State, Myanmar

menglaensis (invalid): China (Yunnan); Type locality: Mengla County, Yunnan Province, China, at an elevation of 700 m elevation.

truongsonicus: Laos: Type locality: Nahin District, Khammouan Province, Laos.

unicolor: Cambodia: Type locality: Kampong Speu Province, Cambodia.  
Reproduction 
TypesLectotype: ZSI 8022, adult male (Figs. 6C and 10 in Poyarkov et al. 2022): collected by T. M. Berdmore, designated by Poyarkov et al. 2022. Other specimens: ZSI.
Holotype. YBU 14124, adult female, collected in June 2014.
Paratypes. YBU 14141 and YBU 14142, two adult males from the same locality as the holotype but collected in July 2012 [menglaensis] 
DiagnosisDiagnosis: Pareas berdmorei differs from other members of the genus Pareas by the following combination of morphological characters: maximal total length of 770 mm; frontal scale hexagonal with its lateral sides parallel to the body axis; the anterior pair of chin shields broader than long; loreal and prefrontal not contacting the eye; generally 1 or 2 preoculars; regularly 2 (rarely 1 or 3) suboculars; generally single postocular (rarely 0 or 2); temporals 3 + 4 or 3 + 3; one to three median vertebral dorsal scale rows slightly enlarged; generally 8 (7–9) infralabials; 15 dorsal scale rows, of them 3–13 scale rows at midbody feebly keeled; 162–187 ventrals; 57–89 subcaudals, all divided; dorsum yellow-brown to orange, dark markings on dorsum variable; thin upper postorbital stripes continue to nape often forming elongated dark markings; iris uniform, color varies from beige to bright reddish-orange (Theobald, 1868b; Bourret, 1934; Taylor, 1965; Ziegler et al., 2006; Yang & Rao, 2008; Le et al., 2014; Pham & Nguyen, 2019; Wang et al., 2020; Poyarkov et al. 2022).


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CommentSynonymy: Poyarkov et al. 2022 revalidated P. bermorei from the synonymy of P. carinatus and synonymized P. Pareas menglaensis with P. berdmorei.

Distribution: see map in Poyarkov et al. 2022: Fig. 1. 
EtymologyTheobald (1868b) named his new species in honor of British naturalist Captain Thomas Matthew Berdmore (1811–1859), who was the collector of the type specimens.

The new subspecies name “truongsonicus” is a Latin toponymic adjective in nominative singular, adopting the masculine gender of the genus name Pareas, and is given in reference to the Truong Son (Annamite) Mountain Range in Vietnam and Laos, where the new subspecies occurs (Poyarkov et al. 2022).

P. menglaensis was named after the type locality, Mengla County, Yunnan, China. 
References
  • DAVID, P., & DEUTI, K. 2022. On the type specimens of Pareas macularius Theobald, 1868 and Pareas berdmorei Theobald, 1868 with the designation of a lectotype for Pareas macularius (Squamata: Serpentes: Pareidae). Zootaxa 5105 (4): 490-500 - get paper here
  • Poyarkov NA, Nguyen TV, Pawangkhanant P, Yushchenko PV, Brakels P, Nguyen LH, Nguyen HN, Suwannapoom C, Orlov N, Vogel G. 2022. An integrative taxonomic revision of slug-eating snakes (Squamata: Pareidae: Pareineae) reveals unprecedented diversity in Indochina. PeerJ 10: e12713 - get paper here
  • Wang P, Che J, Liu Q, Li K, Jin JQ, Jiang K, Shi L, Guo P 2020. A revised taxonomy of Asian snail-eating snakes Pareas (Squamata, Pareidae): evidence from morphological comparison and molecular phylogeny. ZooKeys 939: 45-64 - get paper here
 
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