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Paroedura kloki GLAW, KÖHLER & VENCES, 2018

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common Names 
SynonymParoedura kloki GLAW, KÖHLER & VENCES 2018
Paroedura sp. "Ankarafantsika" — GLAW & VENCES (2007
Paroedura sp. "Ankarafantsika" — GLAW et al. (2014
Paroedura sp. "Ankarafantsika" — MORI et al. (2014
Paroedura karstophila — JACKMAN et al. 2008 
DistributionNW Madagascar (Mahajanga)

Type locality: Sahaovy, camp 0, (15°29'19.9'' S, 49°04'42.6'' E, 607 m a.s.l.), Mahajanga Province, northwestern Madagascar  
TypesHolotype: ZSM 536/2009 (field number ZCMV 11231; Fig. 4B), adult female, collected on 20 June 2009 by J. L. Patton, E. Rajeriarison, T. Rajofiarison, R.-D. Randrianiaina, F. M. Ratsoavina, M. Vences, and D. R. Vieites.
Paratypes. UADBA uncatalogued (ZCMV 11230), adult female, and UADBA uncatalogued (ZCMV 11456), male, both with same data as holotype; ZSM 484/2001 (MV 2001-385) and UADBA uncatalogued (MV 2001- 386), two adult males, from Ankarafantsika National Park, Lac Ravelobe (around 16°18'18.87" S, 46°48'59.87" E, ca. 75 m a.s.l.), Mahajanga Province, northwestern Madagascar, collected on 28 February 2001 by G. Garcia, V. Raherisoa, A. Rasoamamonjinirina, M. Vences and D. R. Vieites. GenBank accession numbers for sequences of the ZSM paratype (Jackman et al. 2008): EF490775 (nd2), EF536229 (nd4), EF490722 (rag1) and EF490696 (pdc). 
DiagnosisDiagnosis. The new species is distinguished from P. androyensis, P. bastardi, P. ibityensis, P. lohatsara, P. maingoka, P. picta, and P. vahiny by having the nostril in contact with the rostral scale; from P. gracilis by absence of a raised vertebral ridge on the body and by shorter forelimbs which are not extending forward beyond tip of snout; from P. masobe by much smaller size (SVL up to 62 mm versus 107 mm), much smaller eyes and absence of a dorsal row of paired spines on the tail; from P. stumpffi by absence of a light line bordering at least the posterior head, absence of a light line along the upper lip, and absence of a brown stripe from the nostril to the neck; from P. tanjaka by smaller size (SVL up to 62 mm versus 102 mm); from P. vazimba by larger size (SVL up to 62 mm versus 49 mm); from P. karstophila by absence of a light marking above the forelimbs and from the two Comoran species P. sanctijohannis and P. stellata by colouration. Paroedura kloki differs from most other species of the P. oviceps clade (P. fasciata, P. homalorhina, P. hordiesi, and P. spelaea, described below) by the presence of spinous tubercles on the proximal whorls of the tail (versus absent); furthermore from P. homalorhina by shorter limbs and a less slender general appearance, and from P. fasciata by slightly larger size (SVL up to 62 mm versus 49 mm) and lack of light transversal bands. Paroedura kloki is morphologically most similar to P. oviceps from which it differs by a higher number of spinous tail whorls at the proximal portion of the tail (ca. 8 versus ca. 4), and by more expressed dorsal tubercles, especially in the neck region (Fig. 7A). In addition, P. kloki differs from all other species in the genus by strong genetic differences (Fig. 1, Table 1), except for P. vahiny for which DNA sequences are not yet available. 
EtymologyThe specific epithet is a patronym honouring Dr. Pascal R. J. Klok (Viersen, Germany) in recognition of his financial support for taxonomic research and nature conservation projects in Madagascar through the BIOPAT initiative ( 
  • GLAW, FRANK; JÖRN KÖHLER & MIGUEL VENCES 2018. Three new species of nocturnal geckos of the Paroedura oviceps clade from xeric environments of Madagascar (Squamata: Gekkonidae). Zootaxa 4433 (2): 305–324 - get paper here
  • Kwet, A. 2019. Liste der im Jahr 2018 neu beschriebenen Reptilien. Elaphe 2019 (3): 52-72
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