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Pelomedusa neumanni PETZOLD, VARGAS-RAMÍREZ, KEHLMAIER, VAMBERGER, BRANCH, DU PREEZ, HOFMEYR, MEYER, SCHLEICHER, ŠIROKÝ & FRITZ, 2014

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Higher TaxaPelomedusidae, Pelomedusoidea, Pleurodira, Testudines (turtles)
Subspecies 
Common NamesE: Neumann’s marsh terrapin 
SynonymPelomedusa neumanni PETZOLD, VARGAS-RAMÍREZ, KEHLMAIER, VAMBERGER, BRANCH, DU PREEZ, HOFMEYR, MEYER, SCHLEICHER, ŠIROKÝ & FRITZ 2014
Pelomedusa neumanni — TTWG 2017: 204
Pelomedusa neumanni — SPAWLS et al. 2018: 51
Pelomedusa neumanni — TTWG 2021 
DistributionKenya, Tanzania, Uganda

Type locality: Kakamega, Kenya, N0°17.04 E34°44.52  
Reproduction 
TypesHolotype: NMP P6V 74974, adult male (National Museum Prague); Paratypes: Museum für Naturkunde, Berlin (ZMB 28356, hatchling, Mangara River, Manyara, Tanzania); Naturhistorisches Museum Wien (NMW 24452, male, Lake Victoria); Zoologisches Forschungsmuseum Alexander Koenig, Bonn (ZFMK 81951, hatchling, Kakamega Forest, Kenya). 
DiagnosisDiagnosis: Medium-sized helmeted terrapins with a known maximum straight carapacial length of 19.4 cm. Pectoral scutes rectangular with wide midseam contact. One large undivided temporal head scale. Two small barbels under chin. Colouration variable; specimens with light horn-coloured carapace and completely yellow plastron and individuals with brownish plastron known to occur; soft parts ventrally lighter than dorsally. Pelomedusa neumanni differs from all other Pelomedusa species except possibly P. gehafie, P. kobe and candidate species B (in which the respective character states are unknown) by the presence of thymine (T) instead of cytosine (C) at position 57, by the presence of thymine (T) instead of cytosine (C) or guanine (G) at position 349, and by the presence of adenine (A) instead of cytosine (C) or guanine (G) at position 353 of the 360-bp-long reference alignment of the 12S rRNA gene (Supporting Information). Pelomedusa neumanni differs from P. gehafie, P. kobe and candidate species B by the presence of thymine (T) instead of cytosine (C) at position 116, by the presence of adenine (A) or thymine (T) instead of cytosine (C) or guanine (G) at position 147, and by the presence of adenine (A) instead of guanine (G) at position 223. In addition, P. neumanni differs from P. gehafie by the presence of adenine (A) instead of guanine (G) at position 256, by the presence of cytosine (C) instead of adenine (A) at position 268, and by the presence of cytosine (C) instead of thymine (T) at position 345. Pelomedusa neumanni differs from P. kobe by the presence of cytosine (C) instead of thymine (T) at position 298. Pelomedusa neumanni differs from candidate species B by the presence of cytosine (C) instead of thymine (T) at positions 103, 268 and 274, by the presence of thymine (T) instead of cytosine (C) at positions 109, 125, 152, 287 and 326, by the presence of cytosine (C) instead of adenine (A) at positions 122, 166 and 303, by a gap instead of thymine (T) at positions 126 and 168, by the presence of adenine (A) instead of guanine (G) at positions 148, 154, 180 and 223, and by the presence of thymine (T) instead of adenine (A) at position 302 [from PETZOLD et al. 2014]. 
CommentDistribution: Map: Petzold et al. 2014

Habitat: freshwater (Ponds, swamps) 
EtymologyDedicated to Oscar Neumann (3 September 1867–17 May 1946) who undertook in the late 19th and early 20th centuries influential expeditions to East Africa, Somalia, Ethiopia, and Sudan. Neumann worked voluntarily for over 40 years in the Museum für Naturkunde, Berlin, but was forced by the Nazi government to leave Germany in 1941. 
References
  • Behangana, M., Ayebare, S., Magala, R., Lukwago, W., Ssekisambu, R. S., Alezuyo, C., ... & Hughes, D. 2023. Establishing baseline data for herpetofauna in western Uganda, an East African biodiversity hotspot: Reptile and amphibian diversity in Uganda. Herpetology Notes, 16, 773-789
  • Behangana, Mathias; Richard Magala, Raymond Katumba, David Ochanda, Stephen Kigoolo, Samuel Mutebi, Daniele Dendi,, Luca Luiselli, and Daniel F. Hughes 2020. Herpetofaunal diversity and community structure in the Murchison Falls-Albert Delta Ramsar site, Uganda: Herpetofaunal diversity. European Journal of Ecology, 6(2)
  • Bour, R. 2008. Global diversity of turtles (Chelonii; Reptilia) in freshwater. Hydrobiologia 595:593–598 - get paper here
  • Fritz, U., T. Mazuch, A. Petzold & H. Prokop 2015. Coloration and pattern of hatchlings of six Pelomedusa species. Salamandra 51 (3): 277-282 - get paper here
  • Fritz, U., Tolley, K. A., Vamberger, M., & Ihlow, F. 2022. Phylogeny and phylogeography of chelonians from sub-Saharan Africa—A review of current knowledge in tribute to Margaretha D. Hofmeyr. Vertebrate Zoology, 72: 951-969 - get paper here
  • Gemel, R.; G. Gassner & S. Schweiger 2019. Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien – 2018. Ann. Naturhist. Mus. Wien, B 121: 33–248
  • Kirchhof S, Wasonga V, Mazuch T, Spawls S, Malonza KP. 2023. An annotated checklist of the herpetofauna of the Sibiloi National Park in northern Kenya based on field surveys. Amphibian & Reptile Conservation 17(1/2) [General Section]: 1–18 (e324) - get paper here
  • Nagy, Z. T., J. Kielgast, M. Moosig, M. Vamberger & U. Fritz 2015. Another candidate species of Pelomedusa (Testudines: Pelomedusidae) from the Democratic Republic of the Congo? Salamandra 51 (2): 212-214 - get paper here
  • PETZOLD, ALICE; MARIO VARGAS-RAMÍREZ, CHRISTIAN KEHLMAIER, MELITA VAMBERGER, WILLIAM R. BRANCH, LOUIS DU PREEZ, MARGARETHA D. HOFMEYR, LEON MEYER, ALFRED SCHLEICHER, PAVEL ŠIROKÝ & UWE FRITZ 2014. A revision of African helmeted terrapins (Testudines: Pelomedusidae: Pelomedusa), with descriptions of six new species. Zootaxa 3795 (5): 523–548 - get paper here
  • Spawls, Stephen; Tomáš Mazuch & Abubakr Mohammad 2023. Handbook of Amphibians and Reptiles of North-east Africa. Bloomsbury, 640 pp. - get paper here
  • Spawls, Steve; Kim Howell, Harald Hinkel, Michele Menegon 2018. Field Guide to East African Reptiles. Bloomsbury, 624 pp. - get paper here
  • TTWG; Anders G.J. Rhodin, John B. iverson, Roger Bour, Uwe Fritz, Arthur GeoRges, H. BRadley Shaffer, and Peter Paul van Dijk 2017. Turtles of the World: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status (8th Ed.). In: Rhodin, A.G.J., Iverson, J.B., van Dijk, P.P., Saumure, R.A., Buhlmann, K.A., Pritchard, P.C.H., and Mittermeier, R.A. (Ed Chelonian Research Monographs 7:1–292. doi: 10.3854/crm.7.checklist.atlas.v8.2017 - get paper here
  • TTWG; Rhodin, A.G.J., Iverson, J.B., Bour, R., Fritz, U., Georges, A., Shaffer, H.B., and van Dijk, P.P. 2021. Turtles of the World: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status (9th Ed.). In: Rhodin, A.G.J., Iverson, J.B., van Dijk, P.P., Stanford, C.B., Goode, E.V., Buhlmann, K.A., and Mittermeier, R.A. (Eds.). Chelonian Research Monographs 8:1–472. doi:10.3854/crm.8.checklist.atlas.v9.2021. - get paper here
 
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