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Pelturagonia cephalum MOCQUARD, 1890

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Mocquard's Eyebrow Lizard 
SynonymPelturagonia cephalum MOCQUARD 1890: 130
Pelturagonia cephalum MOCQUARD 1890: 144
Japalura nigrilabris DE ROOIJ 1915 (non PETERS 1864)
Phoxophrys cephalum — INGER 1960: 223
Phoxophrys cephalum — WERMUTH 1967: 74
Phoxophrys cephalum — BRYGOO 1988: 12
Phoxophrys cephalum — MANTHEY & GROSSMANN 1997: 196
Phoxophrys cephalum — MALKMUS et al. 2002: 248
Phoxophrys cephalum — MANTHEY 2010
Phoxophrys cephalum — MANTHEY & DENZER 2019
Pelturagonia cephalum — HARVEY et al. 2019 
DistributionIndonesia (Borneo, Sabah), Malaysia (Sabah, Sarawak)

Type locality: Kina Balu.  
Reproductionoviparous 
TypesSyntypes: MNHN-RA 1889.0170-0171 
DiagnosisDiagnosis (genus): Medium-sized draconine agamids reaching a SVL of at least 84 mm and distinguished from all other Agamidae by the following combination of characters: (1) tympanum and external auditory meatus absent, extrastapes attaching to skin; (2) head robust, 77.5–98.3% as wide as long, comprising 22.5–28.4% of SVL; (3) rostral vertically divided into two or more scales; (4) prominent tubercular scale positioned below corner of jaw; (5) pair of dorsolateral caudal crests, not separated medially by vertebral crest; (6) dorsal scales heterogenous with larger tubercular scales interspersed among smaller-keeled scales, 81–115 scales around midbody; (7) alveolar portion of premaxilla relatively narrow, usually with single tooth and with premaxillary processes of mandible suturing or fusing over top of it; (8) frontal and parietal extensively sculptured, not containing pineal foramen; (9) conch of quadrate rudimentary, with wide dorsolateral triangular portion; (10) angular process of mandible large, tab-like, and vertically developed. 
CommentSynonymy: BOULENGER (1891) synonymized Pelturagonia cephalum with Japalura nigrilabris. It seems that Mocquard described the (genus and) species twice in two different articles.

Type species: Pelturagonia cephalum MOCQUARD 1890: 130 is the type species of the genus Pelturagonia MOCQUARD 1890. Pelturagonia was considered a synonym of Phoxophrys for decades but has been revalidated by Harvey et al. 2019.

Key: Harvey et al. 2019 provide a dichotomous key and character table (Table 1) to the species of Pelturagonia. 
EtymologyExplaining the derivation of his new name, Mocquard (1890:130) described the distinctive angled aspect of the caudal crests and remarked ‘‘pour cette raison, nous imposons le nom générique de Pelturagonia.’’ Pelturagonia is a feminine Latin noun in the nominative singular derived from the feminine Greek words pelte meaning shield and gonia meaning angle. 
References
  • Ahmad, Norhayati; Eddie Ahmad, Mahathir Ratag, Edward Asrul Alimin Sinon, Brandon Don, Francer Francis, Muhammad Ridzuan Mahmod, Amshari Agimin, Daicus Belabut 2019. Amphibians and Reptiles of Imbak Canyon Study Centre and Batu Timbang Camp. Journal of Tropical Biology and Conservation 16: 25–33
  • Boulenger, G. A. 1891. Remarks on the herpetological fauna of Mount Kina baloo, North Borneo. Ann. Mag. nat. Hist. (6) 7: 341-345 - get paper here
  • Brygoo, E.R. 1988. Les types d’Agamidés (Reptiles, Sauriens) du Muséum national d’Histoire naturelle Catalogue critique. Bull. Mus. Nat. Hist. Nat. 10 (ser. 4) A (3), suppl.: 1-56
  • Das , I. 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, Borneo
  • Das, I. & Yaakob, N. 2007. Status of knowledge of the Malaysian herpetofauna. In Status of biological diversity in Malaysia & threat assessment of plant species in Malaysia. in: L. S. L. Chua, L. G. Kirton & L. G. Saw (eds.), Status of biological diversity in Malaysia & threat assessment of plant species in Malaysia. Forest Research Institute Malaysia, Kepong, pp. 31-81
  • Das, Indraneil 2006. Crocker Range National Park, Sabah, as a refuge for Borneo's montane herpetofauna. Amphibian & Reptile Conservation 4 (1): 3-11 - get paper here
  • Harvey, Michael B., Larson, Thorton R., Jacobs, Justin L., Shaney, Kyle, Streicher, Jeffrey W., Hamidy, Amir, Kurniawan, Nia & Smit, Eric N. 2019. Phoxophrys After 60 Years: Review of Morphology, Phylogeny, Status of Pelturagonia, and a New Species from Southeastern Kalimantan. Herpetological Monographs 33 (1): 71-107 - get paper here
  • Inger, R. F. 1960. A review of the agamid lizards of the genus Phoxophrys HUBRECHT. Copeia 1960 (3): 221 -225. - get paper here
  • Malkmus, R.; Manthey, U.; Vogel, G. Hoffmann, P. & Kosuch, J. 2002. Amphibians and reptiles of Mount Kinabalu (North Borneo). A.R.G. Ganther Verlag, Rugell, 404 pp.
  • Manthey U 2010. Agamid Lizards of Southern Asia. Draconinae 2 -Leiolepidinae. Edition Chimaira, Terralog 7b, Frankfurt, 168 pp.
  • MANTHEY, U. & DENZER, W. 2019. Rare mountain agamas of the Kinabalu Park (Sabah, North Borneo) above 900 m a.s.l. Sauria 41 (1): 2 - 12 - get paper here
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • Mocquard, F. 1890. Diagnoses d'espèces nouvelles de reptiles et de batraciens des iles Bornèo et Palawan. Le Naturaliste, Paris: 144-145. - get paper here
  • Mocquard, F. 1890. Recherches sur la faune herpétologique des îles de Bornèo et de Palawan. Nouv. Arch. Mus. Hist. Nat., Paris, 3 (2): 115-168. - get paper here
 
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