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Philodryas cordata DONNELLY & MYERS, 1991

IUCN Red List - Philodryas cordata - Least Concern, LC
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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common Names 
SynonymPhilodryas cordata DONNELLY & MYERS 1991: 46
Philodryas cordatus — KORNACKER 1999: 123
Philodryas cordata — GRAZZIOTIN et al. 2012
Philodryas cordata — WALLACH et al. 2014: 550
Philodryas cordata — MELO-SAMPAIO et al. 2020 
DistributionVenezuela

Type locality: North side of Cerro Guaiquinima (1030 m elevation), Estado Bolívar, Venezuela.  
Reproductionoviparous 
TypesHolotype: AMNH 136212 (field no. CWM 18974), an adult male 
DiagnosisDiagnosis: A tan Philodryas readily distinguished from all other species occurring north of the Amazon in being tan in life rather than green The species furthermore seems to differ from all other Philodryas in color pattern and penial characters (see DONNELLY & MYERS 1991).


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CommentHabitat: fully arboreal (Harrington et al. 2018). 
EtymologyThe species name is derived from the Latin cordis (genitive of cor, the heart)+ -atus-meaning, in the present connection, "heart-shaped," in allusion to the everted hemipenis (see fig. 33). It is an adjective given in feminine form as demanded by the generic name, which is of feminine gender. 
References
  • Donnelly M A; Myers C W 1991. Herpetological results of the 1990 Venezuelan Expedition to the summit of Cerro Guaiquinima, with new tepui reptiles. American Museum Novitates (3017): 1-54 - get paper here
  • Grazziotin, Felipe G.; Hussam Zaher, Robert W. Murphy, Gustavo Scrocchi, Marco A. Benavides, Ya-Ping Zhang and Sandro L. Bonatto 2012. Molecular phylogeny of the New World Dipsadidae (Serpentes: Colubroidea): a reappraisal. Cladistics DOI: 10.1111/j.1096-0031.2012.00393.x - get paper here
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Kornacker,P.M. 1999. Checklist and key to the snakes of Venezuela. PaKo-Verlag, Rheinbach, Germany, 270 pp.
  • Melo-Sampaio, P. R., Passos, P., Martins, A. R., Moura-leite, J. C., Morato, S. A., Venegas, P. J., Chávez G., Venâncio N.M. & De Souza, M. B. 2020. A phantom on the trees: integrative taxonomy supports a reappraisal of rear-fanged snakes classification (Dipsadidae: Philodryadini). Zoologischer Anzeiger 290: 19-39 [dated 2021 in print version, but 2020 published online] - get paper here
  • Natera-Mumaw, Marco; Luis Felipe Esqueda-González & Manuel Castelaín-Fernández 2015. Atlas Serpientes de Venezuela. Santiago de Chile, Dimacofi Negocios Avanzados S.A., 456 pp. - get paper here
  • RIVAS, GILSON A.; CÉSAR R. MOLINA, GABRIEL N. UGUETO, TITO R. BARROS, CÉSAR L. BAR- RIO-AMORÓS & PHILIPPE J. R. KOK 2012. Reptiles of Venezuela: an updated and commented checklist. Zootaxa 3211: 1–64 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
 
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