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Philodryas arnaldoi (AMARAL, 1932)

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Common NamesE: Arnaldo's Green Racer
Portuguese: Papa-Pinto, Papa-Rato, Parelheira-Clara, Parelheira-do-Mato 
SynonymChlorosoma arnaldoi AMARAL 1932: 100
Philodryas arnaldoi — GRAZZIOTIN et al. 2012
Philodryas arnaldoi — WALLACH et al. 2014: 549
Philodryas arnaldoi — NOGUEIRA et al. 2019
Philodryas arnaldoi — ARREDONDO et al. 2020
Pseudablabes arnaldoi — MELO-SAMPAIO et al. 2020 
DistributionBrazil (Sao Paulo, Parana, Rio Grande do Sul, Santa Catarina)

Type locality: S. Bento, Santa Catarina, Brazil.  
TypesHolotype: IBSP (= IB) 7807, female 
CommentHabitat: fully arboreal (Harrington et al. 2018).

Phylogenetics: MELO-SAMPAIO et al. 2020 tentatively included arnaldoi within Pseudablabes (based on morphology) although they did not have any molecular data. Note that Arredondo et al. 2020 could not replicate the monophyly of Pseudablabes as defined by Melo-Sampaio et al., hence we reverted them to Philodryas. 
EtymologyNamed after Arnaldo França, Amaral's technical assistant. 
  • Amaral,A. do 1932. Contribuicao ao conhecimento dos ofidios do Brasil VI. Uma nova especie de Colubrideo opisthoglypho, do genero Chlorosoma Wagler, 1830. Mem. Inst. Butantan 7: 99-101 + 1 plate - get paper here
  • Arredondo, J. C., Grazziotin, F. G., Manfrini, G. J. S., Rodrigues, M. T. U., Bonatto, S. L., & Zaher, H. E. D. 2020. Molecular phylogeny of the tribe Philodryadini Cope, 1886 (Dipsadidae: Xenodontinae): Rediscovering the diversity of the South American Racers. Papéis Avulsos de Zoologia, 60, e20206053-e20206053 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Gonzalez R. C. et al. 2020. Lista dos Nomes Populares dos Répteis no Brasil – Primeira Versão. Herpetologia Brasileira. 9(2): 121 – 214
  • Grazziotin, Felipe G.; Hussam Zaher, Robert W. Murphy, Gustavo Scrocchi, Marco A. Benavides, Ya-Ping Zhang and Sandro L. Bonatto 2012. Molecular phylogeny of the New World Dipsadidae (Serpentes: Colubroidea): a reappraisal. Cladistics DOI: 10.1111/j.1096-0031.2012.00393.x - get paper here
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Lema, Thales de 1962. Ocorréncia de Philodryas arnaldoi (Amaral, 1932) no estado do Rio Grande do Sul, Brasil (Serpentes, Colubridae). Iheringia, Série Zoologia 22: 1-4 + 2 plates
  • Melo-Sampaio, P. R., Passos, P., Martins, A. R., Moura-leite, J. C., Morato, S. A., Venegas, P. J., Chávez G., Venâncio N.M. & De Souza, M. B. 2020. A phantom on the trees: integrative taxonomy supports a reappraisal of rear-fanged snakes classification (Dipsadidae: Philodryadini). Zoologischer Anzeiger 290: 19-39 [dated 2021 in print version, but 2020 published online] - get paper here
  • Morato, Sérgio Augusto Abrahão; Renato Silveira Bérnils, Julio Cesar de Moura-Leite 2017. RÉPTEIS DE CURITIBA: COLETÂNEA DE REGISTROS Hori Consultoria, Curitiba, Paraná, Brasil, 99 pp.
  • Nogueira, Cristiano C.; Antonio J.S. Argôlo, Vanesa Arzamendia, Josué A. Azevedo,<br />Fausto E. Barbo, Renato S. Bérnils, Bruna E. Bolochio, Marcio Borges-Martins,<br />Marcela Brasil-Godinho, Henrique Braz, Marcus A. Buononato, Diego F. Cisneros 2019. Atlas of Brazilian snakes: verified point-locality maps to mitigate the Wallacean shortfall in a megadiverse snake fauna. South American J. Herp. 14 (Special Issue 1):1-274 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
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