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Philothamnus bequaerti (SCHMIDT, 1923)

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Bequaert's Green Snake 
SynonymChlorophis bequaerti SCHMIDT 1923: 75
Philothamnus irregularis — SCHMIDT 1923: 76 (fide HUGHES 1985)
Philothamnus bequarti — HUGHES 1985 (in error)
Philothamnus bequaerti — BROADLEY 1998
Philothamnus bequaerti — CHIRIO & INEICH 2006
Philothamnus bequaerti — WALLACH et al. 2014: 554
Philothamnus bequaerti — SPAWLS et al. 2018: 487 
DistributionCentral African Republic, Cameroon, Uganda, S Sudan (Jumhūriyyat), Republic of South Sudan (RSS), W Ethiopia, W/N Democratic Republic of the Congo (Zaire)

Type locality: Niangara, Democratic Republic of the Congo  
Reproductionoviparous 
TypesHolotype: AMNH 12080, a 652 mm male (H. Lang and J. P. Chapin, Nov. 1910). 
DiagnosisDiagnosis: Habitus very slender, tail one-third the total length; ventral plates distinctly keeled; anal entire; three labials entering the eye; one anterior temporal; dorsal scales in fifteen rows; ventrals 164-170, subcaudals, 123 (Schmidt 1923).


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CommentHas been confused with P. angolensis by Rasmussen (1991) and is probably absent from the Congo (but present in the Democratic Republic of the Congo!).

Habitat: fully arboreal (Harrington et al. 2018). 
EtymologyNamed after Dr. Joseph Charles Bequaert (1886-1982), a Belgian botanist, entomologist, and malacologist who graduated with a doctorate in botany from the University of Ghent (1906). He later worked for the colonial government in the Belgian Congo (1910-1915). He moved to the USA (1916), becoming a U.S. citizen (1921). He was a Research Assistant, American Museum of Natural History (1917-1922), then worked at Harvard (1923-1956), initially teaching entomology at Harvard Medical School and finally becoming Professor of Zoology, Museum of Comparative Zoology. See Beolens et al. 2011 for more details. 
References
  • Behangana, Mathias; Richard Magala, Raymond Katumba, David Ochanda, Stephen Kigoolo, Samuel Mutebi, Daniele Dendi,, Luca Luiselli, and Daniel F. Hughes 2020. Herpetofaunal diversity and community structure in the Murchison Falls-Albert Delta Ramsar site, Uganda: Herpetofaunal diversity. European Journal of Ecology, 6(2)
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Broadley, D.G. 1998. The reptilian fauna of the Democratic Republic of the Congo (Congo-Kinshasa). In: Schmidt, K.P. and Noble, G.K., Contributions to the Herpetology of the Belgian Congo... [reprint of the 1919 and 1923 papers]. SSAR Facsimile reprints in Herpetology, 780 pp.
  • Chippaux, Jean-Philippe & Kate Jackson 2019. Snakes of Central and Western Africa. Johns Hopkins University Press, 448 pp. [detaileld review in HR 51 (1): 161] - get paper here
  • Chirio, L. & Lebreton, M. 2007. Atlas des reptiles du Cameroun. MNHN, IRD, Paris 688 pp.
  • Chirio, Laurent and Ivan Ineich 2006. Biogeography of the reptiles of the Central African Republic. African Journal of Herpetology 55(1):23-59. - get paper here
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Hughes, B. 1985. Progress on a taxonomic revision of African green tree snakes (Philothamnus spp.). In: Schuchmann, K.L. (ed.) Proceedings of the International Symposium on African Vertebrates. ZFMK, Bonn, pp. 511-530
  • JACOBSEN, N.H.G. 2009. A contribution to the herpetofauna of the Passendro Area, Central African Republic. African Herp News (47): 2-20 - get paper here
  • Largen, M.J.; Spawls, S. 2010. Amphibians and Reptiles of Ethiopia and Eritrea. Edition Chimaira, Frankfurt, 694 pp.
  • Largen,M.J. & Rasmussen,J.B. 1993. Catalogue of the snakes of Ethiopia (Reptilia Serpentes), including identification keys. Tropical Zoology 6: 313-434 - get paper here
  • Rasmussen,J.B. 1991. Snakes (Reptilia: Serpentes) from the Kouilou River basin, including a tentative key of the snakes of the République du Congo. Tauraco Report 4: 175-188
  • Schmidt, K. P. 1923. Contributions to the herpetology of the Belgian Congo based on the collection of the American Museum Congo Expedition, 1909-1915. Part II. Snakes, with field notes by Herbert Lang and James P. Chapin. Bull. Amer. Mus. nat. Hist. 49 (1): 1-146 - get paper here
  • Spawls, S.; Howell, K.; Drewes, R.C. & Ashe, J. 2002. A field guide to the reptiles of East Africa. Academic Press, 543 pp. [reviews in HR 34: 396 and Afr. J. Herp. 51; 147] - get paper here
  • Spawls, Stephen; Tomáš Mazuch & Abubakr Mohammad 2023. Handbook of Amphibians and Reptiles of North-east Africa. Bloomsbury, 640 pp. - get paper here
  • Spawls, Steve; Kim Howell, Harald Hinkel, Michele Menegon 2018. Field Guide to East African Reptiles. Bloomsbury, 624 pp. - get paper here
  • Trape, J.-F. 2023. Guide des serpents d’Afrique occidentale, centrale et d’Afrique du Nord. IRD Éditions, Marseille, 896 pp.
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
 
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