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Phrynocephalus versicolor STRAUCH, 1876

IUCN Red List - Phrynocephalus versicolor - Least Concern, LC
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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards)
SubspeciesPhrynocephalus versicolor doriai BEDRIAGA 1909
Phrynocephalus versicolor siebenrocki BEDRIAGA 1909
Phrynocephalus versicolor versicolor STRAUCH 1876 
Common NamesE: Tuvan Toad-headed Agama
Russian: Пестрая круглоголовка
Chinese: 变色沙蜥 
SynonymPhrynocephalus versicolor STRAUCH 1876: 18
Phrynocephalus versicolor — BOULENGER 1885: 374
Phrynocephalus guentheri BEDRIAGA 1906: 168
Phrynocephalus blanfordi BEDRIAGA 1907
Phrynocephalus ciliaris BEDRIAGA 1907
Phrynocephalus versicolor var. Doriai BEDRIAGA 1909: 332
Phrynocephalus versicolor var. Bogdanowi BEDRIAGA 1909: 334
Phrynocephalus helioscopus var. levis KASHTCHENKO 1909: 122
Phrynocephalus Potanini BEDRIAGA 1909: 389
Phrynocephalus pewzowi
Phrynocephalus versicolor versicolor — NIKOLSKY 1915: 188
Phrynocephalus versicolor doriai — NIKOLSKY 1915: 190
Phrynocephalus versicolor bogdanowi — NIKOLSKY 1915: 191
Phrynocephalus potanini — STEJNEGER 1926: 42
Phrynocephalus rostralis ZAREVSKY 1930
Phrynocephalus parvulus ZAREVSKY 1927 (fide POPE 1935)
Phrynocephalus stejnegeri BANNIKOV 1958: 78 (nomen nudum)
Phrynocephalus versicolor — WERMUTH 1967: 89
Phrynocephalus immaculatus ZHAO 1995: 47
Phrynocephalus versicolor — WANG et al. 2003
Phrynocephalus (Phrynocephalus) versicolor versicolor — BARABANOV & ANANJEVA 2007
Phrynocephalus immaculatus — ANANJEVA 2011
Phrynocephalus versicolor — DUNAYEV et al. 2021
Phrynocephalus przewalskii — SOLOVYEVA et al. 2023: 26

Phrynocephalus versicolor siebenrocki BEDRIAGA 1909
Phrynocephalus versicolor var. siebenrocki BEDRIAGA 1909: 232
Phrynocephalus versicolor var. Siebenrocki — BEDRIAGA 1909: 333
Phrynocephalus versicolor siebenrocki — NIKOLSKY 1915: 191
Phrynocephalus versicolor siebenrocki — MILTO & BARABANOV 2012: 165 
DistributionW China (Xinjiang, Gansu, Ningxia, Nei Mongol = Inner Mongolia), (Siberia), SE Russia (Tuva), Mongolia

doriai: Chian (Zinjiang); Terra typica restricta: "See Ebi-nor" [= Ebinur Lake, Xinjiang Uygur Autonomous Region, China].

Type locality: Alashan Desert, N China  
Reproductionoviparous 
TypesLectotype: ZISP 3929.1 (designated by Peters, 1984: 36)
Lectotype: ZISP 5117.1 (designated by Peters, 1984). “See Ebinor” [= Ebinur lake, Xinjiang Uygur Autonomous Region, China]. Leg: N. M. Przewalsky, 1878; however, Barabanov & Ananjeva 2007: 20 state that there is no type [doriai]
Holotype: ZISP 5156, “zwischen Metschin-Ola und der Stadt Barkul” [between Mieh-Ch’eng-O-La Shan (about 30 km northeast of Barkol) and Barkol, approx. 43°55’ N 93°00 E, Barkol county, Xinjiang Uygur Autonomous Region, China]. Leg: G. N. Potanin, 1878 [siebenrocki] 
DiagnosisUnfortunately we had to temporarily remove additional information as this was scraped by multiple AI companies who sell that data to their customers. These details, e.g. detailed descriptions or comparisons (about 3851 characters), are available for collaborators and contributors. Please contact us for details. 
CommentSubspecies: Wang & Fu (2004) suggested to apply the name P. versicolor only to the populations in the northwestern part of its distribution area. Populations from Tengger Desert, Ordos highland, and further east include P. frontalis, P. przewalskii, and P. versicolor and should be called P. przewalskii. Some populations may contain a new species that remains to be described. Guo & Wang (2007) recognized populations in Kuytun, Xinjiang Uygur Autonomous Region as P. guttatus instead of P. versicolor. Dunayev et al. 2021 elevated hispidus and kulagini to full species status and thus removed them from versicolor, but they did not comment on either doriai or siebenrocki. Ganbold et al. 2022 considered kulagini and hispidus as subspecies of versicolor, but showed in their phylogenetic analysis that they are paraphyletic with versicolor, hence we keep them as valid species (note, however, that their phylogeny is only based on a 522 bp fragment of ND2).

Synonymy partly after BARABANOV & ANANJEVA (2007) who consider doriai and hispidus as synonyms of versicolor (and thus recognize only 2 subspecies: versicolor and kulagini), but see subspecies above. 
References
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