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Phrynocephalus sakoi MELNIKOV, MELNIKOVA, NAZAROV, AL-JOHANY & ANANJEVA, 2015

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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common Names 
SynonymPhrynocephalus sakoi MELNIKOV, MELNIKOVA, NAZAROV, AL-JOHANY & ANANJEVA 2015
Phrynocephalus sakoi — SOLOVYEVA et al. 2023 
DistributionNE Oman

Type locality: Al Sharqiyah Sands, Filim vicinity (20°41’19.7’’ N, 58°17’41.6’’ E), northeastern Oman.  
Reproduction 
TypesHolotype: ZISP 28705, adult male (Fig. 4 in Melnikov et al. 2015). Paratypes. ZISP 28706 – 28710, ZISP 27089, CAS 251008, 251022, 251023 (Fig. 4 in Melnikov et al. 2015) 
DiagnosisDiagnosis: A large and “heavy” Phrynocephalus species with tail longer than body in both sexes; five-star shape bright orange pattern on head in females and black in males, two bright orange stripes on the neck in females and black in males, two longitudinal rows of six bright orange (salmon) patches on the dorsum in females and black in males; with white undertail coloration without bands in both sexes with black distal third in males and black quarter in females (two bands) and no bands in white proximal part.


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Comment 
EtymologyThe species has been “dedicated to the memory of our close friend and colleague Sako Tuniyev [1983-2015], who tragically passed away on January 5, 2015.” 
References
  • Carranza S, Xipell M, Tarroso P, Gardner A, Arnold EN, Robinson MD, et al. 2018. Diversity, distribution and conservation of the terrestrial reptiles of Oman (Sauropsida, Squamata). PLoS One 13 (2): e0190389 - get paper here
  • Carranza, Salvador; Johannes Els; Bernat Burriel-Carranza 2021. A field guide to the reptiles of Oman. Madrid : Consejo Superior de Investigaciones Científicas, 223 pp. [review in HR 53 (3): 531] - get paper here
  • Melnikov, Daniel; Ekaterina Melnikova, Roman Nazarov, Awadh Al-Johany, and Natalia B. Ananjeva 2015. A NEW SPECIES OF Phrynocephalus (AGAMIDAE, SAURIA) FROM AL SHARQIYAH SANDS, NORTHEASTERN OMAN, DEDICATED TO THE MEMORY OF SAKO TUNIYEV (1983 – 2015). Russian Journal of Herpetology 22 (4): 301 – 309
  • Solovyeva, E. N., Dunayev, E. A., Nazarov, R. A., Bondarenko, D. A., & Poyarkov, N. A. 2023. COI-Barcoding and Species Delimitation Assessment of Toad-Headed Agamas of the Genus Phrynocephalus (Agamidae, Squamata) Reveal Unrecognized Diversity in Central Eurasia. Diversity 2023, 15, x - get paper here
  • Tuniyev B.S. 2015. In memory of Sako B. Tuniev (1983–2015). Current Studies in Herpetology 15 (1/2): 89–94 - get paper here
 
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