Phrynocephalus axillaris BLANFORD, 1875
Can you confirm these amateur observations of Phrynocephalus axillaris?
|Higher Taxa||Agamidae (Agaminae), Sauria, Iguania, Squamata (lizards)|
|Common Names||E: Yarkand toad-headed agama|
|Synonym||Phrynocephalus axillaris BLANFORD 1875: 192|
Phrynocephalus axillaris — BOULENGER 1885: 378
Phrynocephalus axillaris var. idae BEDRIAGA 1907: 168
Phrynocephalus axillaris var. murielis BEDRIAGA 1907
Phrynocephalus axillaris var. ericae BEDRIAGA 1907
Phrynocephalus koslowi BEDRIAGA 1906: 162
Phrynocephalus kozlovi BEDRIAGA 1906 (fide ANANJEVA 2011)
Phrynocephalus axillaris var. Klemenzi BEDRIAGA 1907
Phrynocephalus axillaris var. maculata BEDRIAGA 1907 (nec ANDERSON 1872)
Phrynocephalus axillaris — ZUGMAYER 1909: 498
Phrynocephalus ludovici MOCQUARD 1910
Phrynocephalus axillaris — WERMUTH 1967: 76
Phrynocephalus nasatus GOLUBEV & DUNAYEV 1995
Phrynocephalus axillaris — WANG et al. 2003
Phrynocephalus (Phrynocephalus) axillaris — BARABANOV & ANANJEVA 2007
Phrynocephalus nasatus — DUNAYEV 2020
|Distribution||C Asia (E Turkestan, Dsungarei, Mongolia, Tibet)|
China (Xinjiang, Gansu)
Type locality: Eastern Turkestan, in the plains.
nasatus: W China; Type locality: between the mountain pass of Topa-Duvan (Topa-Bell, Karaduvan Ridge, 41° 11’ N, 80° 48’ E, and the town of Aksu (41° 11’ N, 80° 14’ E), Kashgaria, western China.
|Types||Syntypes: ZSI 3612-18, 3620, 3622-23, 3625-38, 3832-34, 4839-40. Most types of the various synonyms are in ZISP.|
|Diagnosis||Diagnosis (nasatus): distinctly bulbous nasal shields fonn two kidney-shaped growths on the snout. The nostril opens in the lower nasal shield and is directed sideways from the longitudinal axis and down so that it can be seen only from below. The space between the nostrils is very wide and exceeds four to five times the space from the nostril to the edge of the supralabial fold. L.IL.cd. of males is 0.70-0.72, of the female and juvenile specimens 0.79-0.80. The front part of the snout is protuberant and breaks rather sharply at its end. A row of one to four enlarged shields (the shields of young specimens are less distinct) is situated along the center of the snout from the frontal curve to the edge of the upper lip fold near the rostral shield (Golubev & Dunayev 1995).|
Holotype description. Medium-sized toad agama with a relatively long tail, which gets thi1111erra ther sharply towards the end. Except for the scales of the upper parts of the thighs and the distal third of the tail, which have some traces of ridges, the rest of the scales are smooth. The cap is round in shape, its back is not raised. It is covered by scales which are only slightly protuberant and larger in the front. The sinciput eye is surrounded by one or two rows of small scales, behind which in the back paii of the cap are several large scales, odd in shape. The regular scales on the back of the animal get smaller in size toward the sides of the body, and the darker patches of the pattern have thicker thornshaped scales. The lighter scales scattered on the back are also thick, but they are not thorn-shaped. The chin shield is joined from the right and from the left by a row of three to six lower jaw shields that gradually get smaller in size and which (the first pair excepted) do not touch the shields of the lower lip. The rest of the throat scales are slightly protuberant and more or less uniform. The scales on the bottom part of the body ai e flat, tile-like in appearance, with faintly recognizable thorns in the breast area. On the right and left sides at the beginning of the tail there are a number of polygonal thick scales fom1ing small but well visible thorns. The subdigital lamellae of the fourth toes of the hind limbs have two (or more rarely three) quite low ribs of the same size, each ending as a little thorn. The side of that toe that touches the fifth toe has a row of small sharpened fringes. The opposite side of the toe is devoid of them (Golubev & Dunayev 1995).
For a more detailed description and table comparing nasatus and axillaris, see Dunayev 2020.
|Comment||Synonymy after WERMUTH 1967 and BARABANOV & ANAJEVA 2007. Dunayev 2020 “revalidated” P. nasatus but his description is unsatisfactory, e.g. because there were no localities of specimens given (neither in a list nor on a map). The characters pointed out to diagnose nasatus from axillaris are not shown (and if they are shown, as in Fig. 6, it’s not even clear which species is shown), hence we retain nasatus in axillaris for the time being. Notably, GenBank has 5 sequences of “nasatus” as of Feb. 2022 (citing Dunayev 2020), but oddly, the paper does not even mention those sequences or any other molecular analysis.|
Type locality (nasatus): the locality, as described appears to be in a heavily degraded agricultural area now.
NCBI taxonID: 2802824 [nasatus]