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Phrynocephalus lutensis KAMALI & ANDERSON, 2015

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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards)
Common NamesE: Lut Desert Toad Headed Agama 
SynonymPhrynocephalus lutensis KAMALI & ANDERSON 2015
Phrynocephalus lutensis — NASRABADI et al. 2017 
DistributionIran (Kerman; probably Khorasan, Sistan, Baluchistan)

Type locality: Rig-e Yalan, Dasht-e Lut (Mega Dune, Lut Desert), Kerman Province, near the junction of Kerman, South Khorasan and Sistan & Baluchistan Provinces. 30° 27’ N, 59° 21’ E. 963m  
TypesHolotype: ZISP 28014, Adult male (Zoological Institute, St. Petersburg). Collected by Mehrdad Ghazvinian 11 March 2014. Paratypes. Adult male, ZISP 28015; Adult female, ZISP 28016, Same data as holotype. 
DiagnosisDiagnosis. No upraised swollen scales on dorsum; no cutaneous fold at angle of mouth; no fringe of scales on posterior border of thigh and sides of base of tail; side of head and neck without projecting fringe-like scales; prominent fold on lateral sides of body, separating dorsal and ventral scales; dorsal scales subequal, smooth in general appearance, with upturned mucros when viewed from the side under magnification, homogeneous, not keeled; no enlarged scales along flanks; nasals not in contact and separated by 1–3 small scales; tail 106–119 percent of snout-vent length; no spinose scales on neck or back of head; both sides of third and fourth toes and one side of second toe strongly fringed; suborbital scales small and equal in size and never elongate; nostril not visible in entirety when head viewed from side; supralabial scales separated from eye by 3–4 rows of scales; about 30 lamellae under fourth toe; width of space between nostrils less than one half of distance between nostril and preocular ridge; about 30 scales from eye to eye across head. Snout-vent length (mm) 65–75, tail 69–89 (tail = 106–119% body length). 
CommentAbundance: only known from its original description (Meiri et al. 2017).

Phylogenetics: Ebrahimipour et al. 2021 found populations of P. maculatus to be paraphyletic, especially including P. lutensis. 
EtymologyNamed after the type locality. 
  • Ebrahimipour, F., Rastegar-Pouyani, N., Rastegar-Pouyani, E., Hosseinian Yousefkhani, S. S., & Kamali, K. 2021. Molecular phylogenetic relationships within the Phrynocephalus maculatus-arabicus species complex (Sauria: Agamidae) in Iran. Journal of Zoological Systematics and Evolutionary Research, 59, 1583–1588
  • KAMALI, KAMRAN & STEVEN C. ANDERSON 2015. A New Iranian Phrynocephalus (Reptilia: Squamata: Agamidae) from the hottest place on earth and a key to the genus Phrynocephalus in southwestern Asia and Arabia. Zootaxa 3904 (2): 249–260 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • NASRABADI, REZA; NASRULLAH RASTEGAR-POUYANI, ESKANDAR RASTEGAR-POUYANI, AHMAD GHARZI 2017. A revised key to the lizards of Iran (Reptilia: Squamata: Lacertilia). Zootaxa 4227 (3): 431–443 - get paper here
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