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Phrynocephalus luteoguttatus BOULENGER, 1887

IUCN Red List - Phrynocephalus luteoguttatus - Least Concern, LC
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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Yellow-speckled toad-headed agama 
SynonymPhrynocephalus luteoguttatus BOULENGER 1887: 497
Phrynocephalus luteo-guttatus — ALCOCK & FINN 1897
Phrynocephalus luteoguttatus — SMITH 1935: 235
Phrynocephalus luteoguttatus — WERMUTH 1967: 83
Phrynocephalus luteoguttatus — KHAN 1980: 132
Phrynocephalus luteoguttatus — KHAN 1999
Phrynocephalus luteoguttatus — ANDERSON 1999: 89
Phrynocephalus (Phrynocephalus) luteoguttatus — BARABANOV & ANANJEVA 2007
Phrynocephalus luteoguttatus — SINDACO & JEREMČENKO 2008
Phrynocephalus (Microphrynocephalus) luteoguttatus — SOLOVYEVA et al. 2014 
DistributionSW Afghanistan, SW Pakistan (Baluchistan)

Type locality: between Nushki and Helmand (border between Afghanistan and Baluchistan).  
Reproductionoviparous 
TypesSyntypes: BMNH 1946.8.28.44-49 (was BMNH 86.9.21.62-64) and BMNH 1946.8.28.36-38 (was BMNH 86.9.21.69-71); ZISP 7363 (was BMNH cnu). 
DiagnosisDiagnosis: About 30 scales from eye to eye across head; nasals in contact or partly
separated; nostrils directed more or less directly forward; dorsal scales heterogeneous, enlarged scales strongly keeled, imbricate, flat, not tuberculate, posterior border not sharply upturned; sides of back of head and neck with long, flat, upturned fringe-like scales; both sides of fourth toe with long, well-developed fringes. Tail 85-102 percent of snout-vent length (Anderson 1999: 89).

Color pattern: Dorsum yellowish brown to pale buff, with black dots and grayish reticulum enclosing pale yellow or golden spots; sides sometimes blackish, a blackish streak along outer side ofleg present or absent; venter whitish (pink in life). Tip of tail with black crossbars or asymmetrical spots on lower surface (juveniles), or entire tip (more than distal third) black (adults). Boulenger (1889a, pI. 8, fig. 4) shows colors in life. In preserved specimens (from Afghanistan) a pink tinge is still visible in light areas on under side of tail (Anderson 1999: 89). 
CommentNot in India according to KHAN 1999. DAS (1997) mentions it but presently doubtful for India. Not in Iran fide ŠMÍD et al. 2014 (lizard checklist Iran). 
References
  • Alcock, A. W. & FINN,F. 1897. An account of the Reptilia collected by Dr. F.P. Maynard, Captain A.H. McMahon, C.I.E., and the members of the Afghan-Baluch Boundary Comission of 1896. J. Asiat. Soc. Bengal 65: 550-566 [1896] - get paper here
  • Anderson, Steven C 1999. The lizards of Iran. Contributions to Herpetology Volume 15, Society for the Study of Amphibians and Reptiles, Saint Louis, Missouri: i-vii, 1-442 [review in Copeia 2000 (4): 1144] - get paper here
  • BARABANOV, A.V. & N.B. ANANJEVA 2007. Catalogue of the available scientific species-group names for lizards of the genus Phrynocephalus Kaup, 1825 (Reptilia, Sauria, Agamidae). Zootaxa 1399: 1-56 - get paper here
  • Boulenger, G. A. 1887. Catalogue of the lizards in the British Museum (Nat. Hist.) III. Lacertidae, Gerrhosauridae, Scincidae, Anelytropsidae, Dibamidae, Chamaeleontidae. London: 575 pp. - get paper here
  • KAMALI, KAMRAN & STEVEN C. ANDERSON 2015. A New Iranian Phrynocephalus (Reptilia: Squamata: Agamidae) from the hottest place on earth and a key to the genus Phrynocephalus in southwestern Asia and Arabia. Zootaxa 3904 (2): 249–260 - get paper here
  • Khan, M. S. 1999. A checklist and key to the phrynocephalid lizards of Pakistan, with ethnological notes (Squamata: Agamidae). Pakistan J. Zool. 31 (1): 17-24 - get paper here
  • Khan, M.S. 1980. Affinities and zoogeography of herptiles of Pakistan. Biologia, 26(1-2): 113-171
  • Khan, M.S. 2003. CHECKLIST AND KEY TO THE LIZARDS OF PAKISTAN. Pakistan J. Zool. Suppl. Ser. (1): 1-25 - get paper here
  • Leviton, Alan E. 1959. Report on a collection of reptiles from Afghanistan. Proc. Cal. Acad. Sci. 29 (12): 445-463 - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Smith, M.A. 1935. The fauna of British India, including Ceylon and Burma. Reptiles and Amphibia, Vol. II. Sauria. Taylor and Francis, London, 440 pp.
  • Solovyeva, E. N.; N. A. Poyarkov, E. A. Dunayev, R. A. Nazarov, V. S. Lebedev, and A. A. Bannikova 2014. Phylogenetic Relationships and Subgeneric Taxonomy of Toad-headed Agamas Phrynocephalus (Reptilia, Squamata, Agamidae) as Determined by Mitochondrial DNA Sequencing. Doklady Biological Sciences 455: 119–124. - get paper here
  • Werner,F. 1895. Über eine Sammlung von Reptilien aus Persien, Mesopotamien und Arabien. Verh. Zool.-Bot. Ges. Wien 45: 13-22 - get paper here
 
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