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Phrynocephalus luteoguttatus BOULENGER, 1887

IUCN Red List - Phrynocephalus luteoguttatus - Least Concern, LC
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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Yellow-speckled toad-headed agama 
SynonymPhrynocephalus luteoguttatus BOULENGER 1887: 497
Phrynocephalus luteo-guttatus — ALCOCK & FINN 1897
Phrynocephalus luteoguttatus — SMITH 1935: 235
Phrynocephalus luteoguttatus — WERMUTH 1967: 83
Phrynocephalus luteoguttatus — KHAN 1980: 132
Phrynocephalus luteoguttatus — KHAN 1999
Phrynocephalus luteoguttatus — ANDERSON 1999: 89
Phrynocephalus (Phrynocephalus) luteoguttatus — BARABANOV & ANANJEVA 2007
Phrynocephalus luteoguttatus — SINDACO & JEREMČENKO 2008
Phrynocephalus (Microphrynocephalus) luteoguttatus — SOLOVYEVA et al. 2014
Phrynocephalus (Microphrynocephalus) luteoguttatus — SOLOVYEVA et al. 2023 
DistributionSW Afghanistan, SW Pakistan (Baluchistan)

Type locality: between Nushki and Helmand (border between Afghanistan and Baluchistan).  
Reproductionoviparous 
TypesSyntypes: BMNH 1946.8.28.44-49 (was BMNH 86.9.21.62-64) and BMNH 1946.8.28.36-38 (was BMNH 86.9.21.69-71); ZISP 7363 (was BMNH cnu). 
DiagnosisDiagnosis: About 30 scales from eye to eye across head; nasals in contact or partly
separated; nostrils directed more or less directly forward; dorsal scales heterogeneous, enlarged scales strongly keeled, imbricate, flat, not tuberculate, posterior border not sharply upturned; sides of back of head and neck with long, flat, upturned fringe-like scales; both sides of fourth toe with long, well-developed fringes. Tail 85-102 percent of snout-vent length (Anderson 1999: 89).


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CommentNot in India according to KHAN 1999. DAS (1997) mentions it but presently doubtful for India. Not in Iran fide ŠMÍD et al. 2014 (lizard checklist Iran). 
References
  • Alcock, A. W. & FINN,F. 1897. An account of the Reptilia collected by Dr. F.P. Maynard, Captain A.H. McMahon, C.I.E., and the members of the Afghan-Baluch Boundary Comission of 1896. J. Asiat. Soc. Bengal 65: 550-566 [1896] - get paper here
  • Anderson, Steven C 1999. The lizards of Iran. Contributions to Herpetology Volume 15, Society for the Study of Amphibians and Reptiles, Saint Louis, Missouri: i-vii, 1-442 [review in Copeia 2000 (4): 1144] - get paper here
  • BARABANOV, A.V. & N.B. ANANJEVA 2007. Catalogue of the available scientific species-group names for lizards of the genus Phrynocephalus Kaup, 1825 (Reptilia, Sauria, Agamidae). Zootaxa 1399: 1-56 - get paper here
  • Boulenger, G. A. 1887. Catalogue of the lizards in the British Museum (Nat. Hist.) III. Lacertidae, Gerrhosauridae, Scincidae, Anelytropsidae, Dibamidae, Chamaeleontidae. London: 575 pp. - get paper here
  • KAMALI, KAMRAN & STEVEN C. ANDERSON 2015. A New Iranian Phrynocephalus (Reptilia: Squamata: Agamidae) from the hottest place on earth and a key to the genus Phrynocephalus in southwestern Asia and Arabia. Zootaxa 3904 (2): 249–260 - get paper here
  • Khan, M. S. 1999. A checklist and key to the phrynocephalid lizards of Pakistan, with ethnological notes (Squamata: Agamidae). Pakistan J. Zool. 31 (1): 17-24 - get paper here
  • Khan, M.S. 1980. Affinities and zoogeography of herptiles of Pakistan. Biologia, 26(1-2): 113-171
  • Khan, M.S. 2003. CHECKLIST AND KEY TO THE LIZARDS OF PAKISTAN. Pakistan J. Zool. Suppl. Ser. (1): 1-25 - get paper here
  • Leviton, Alan E. 1959. Report on a collection of reptiles from Afghanistan. Proc. Cal. Acad. Sci. 29 (12): 445-463 - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Smith, M.A. 1935. The fauna of British India, including Ceylon and Burma. Reptiles and Amphibia, Vol. II. Sauria. Taylor and Francis, London, 440 pp.
  • Solovyeva, E. N., Dunayev, E. A., Nazarov, R. A., Bondarenko, D. A., & Poyarkov, N. A. 2023. COI-Barcoding and Species Delimitation Assessment of Toad-Headed Agamas of the Genus Phrynocephalus (Agamidae, Squamata) Reveal Unrecognized Diversity in Central Eurasia. Diversity 2023, 15, x - get paper here
  • Solovyeva, E. N.; N. A. Poyarkov, E. A. Dunayev, R. A. Nazarov, V. S. Lebedev, and A. A. Bannikova 2014. Phylogenetic Relationships and Subgeneric Taxonomy of Toad-headed Agamas Phrynocephalus (Reptilia, Squamata, Agamidae) as Determined by Mitochondrial DNA Sequencing. Doklady Biological Sciences 455: 119–124. - get paper here
  • Werner,F. 1896. Über eine Sammlung von Reptilien aus Persien, Mesopotamien und Arabien. Verh. Zool.-Bot. Ges. Wien 45: 13-22 [1895] - get paper here
 
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