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Phrynosoma goodei STEJNEGER, 1893

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Higher TaxaPhrynosomatidae, Phrynosomatinae, Phrynosomatini; Iguania, Sauria, Squamata (lizards)
Common NamesE: Sonoran Horned Lizard
G: Sonora-Wüstenkrötenechse
S: Camaleón de Sonora 
SynonymPhrynosoma platyrhinos goodei STEJNEGER 1893: 191
Anota goodei — COPE 1896: 834
Phrynosoma goodei — MULCAHY et al. 2006
Phrynosoma goodei — COLLINS & TAGGART 2009
Phrynosoma (Doliosaurus) goodei — CROTHER et al. 2012 
DistributionUSA (SW Arizona), Mexico (NW Sonora)

Type locality: "Coast deserts of the state of Sonora, Mexico," restricted to Puerto Libertad by SMITH & TAYLOR 1950.  
TypesHolotype: USNM 8567a 
CommentMULCAHY et al. 2006 elevated Phrynosoma platyrhinos goodei to full species status based on mtDNA sequences. MULCAHY et al. 2006 also presented evidence for goodei x mcalli hybrids.

Group: Belongs to the Doliosaurus clade fide LEACHE & MCGUIRE 2006. 
EtymologyNamed after George Brown Goode (1851-1896), American ichthyologist, assistant to Baird, and Assistant Secretary ofthe Smithsonian from 1872. 
  • Aguilar-Morales, Cecilia and Thomas R. Van Devender 2018. Horned Lizards (Phrynosoma) of Sonora, Mexico: Distribution and Ecology. Sonoran Herpetologist 31 (3): 40-50 - get paper here
  • Baird, S.F. and Girard,C. 1852. Characteristics of some new reptiles in the Museum of the Smithsonian Institution. Proc. Acad. Nat. Sci. Philadelphia 6: 68-70 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Bezy, R. L., P. C. Rosen, T. R. Van Devender, and E. F. Enderson. 2017. Southern distributional limits of the Sonoran Desert herpetofauna along the mainland coast of northwestern Mexico. Mesoamerican Herpetology 4(1): 138–167 - get paper here
  • Bonetti, Mathilde 2002. 100 Sauri. Mondadori (Milano), 192 pp. - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Bruekers, Jaco and Peek, Ron 2004. Veldwaarnemingen aan de zuidelijke woestijn-Padhagedis, Phrynosoma platyrhinos calidiarum (COPE 1896) in Nevada en Zuid-Californië. Lacerta 62 (3): 106-112 [erratum in 62 (4): 174] - get paper here
  • Burt, Charles E. 1933. Some lizards from the Great Basin of the West and adjacent areas, with comments on the status of various forms. American Midland Naturalist 14: 228-250 - get paper here
  • Cope, E.D. 1896. On two new species of lizards from Southern California. American Naturalist 30: 833-836 - get paper here
  • Cope, E.D. 1900. The crocodilians, lizards and snakes of North America. Ann. Rep. U.S. Natl. Mus. 1898: 153-1270 - get paper here
  • Crother, B. I. (ed.) 2012. Standard Common and Current Scientific Names for North American Amphibians, Turtles, Reptiles, and Crocodilians, Seventh Edition. Herpetological Circular 39: 1-92
  • Gentry, A.F. 1885. A review of the genus Phrynosoma. Proc. Acad. Nat. Sci. Philad. (ser. 3) 37: 138-148 - get paper here
  • Girard, Charles F. 1858. United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, Under the command of Charles Wilkes, U.S.N. Vol. 20. Herpetology. C. Sherman & Son, Philadelphia, xv, 492 pages [see note in Zhao and Adler 1993: 369] - get paper here
  • Jones, L.L. & Lovich, R.E. 2009. Lizards of the American Southwest. A photographic field guide. Rio Nuevo Publishers, Tucson, AZ, 568 pp. [review in Reptilia 86: 84] - get paper here
  • Klauber, Laurence M. 1935. The status of the Sonoran horned toad, Phrynosoma goodei Stejneger. Copeia 1935 (4): 178-179 - get paper here
  • Lara-Resendiz, Rafael Alejandro, Tereza Jezkova, Philip C. Rosen and Fausto Roberto Mendez-de la Cruz. 2014. Thermoregulation during the summer season in the Goode's Horned Lizard Phrynosoma goodei (Iguania: Phrynosomatidae) in Sonoran Desert. Amphibia-Reptilia 35 (2): 161-172 - get paper here
  • Leaché, Adam D. and Jimmy A. McGuire 2006. Phylogenetic relationships of horned lizards (Phrynosoma ) based on nuclear and mitochondrial data: Evidence for a misleading mitochondrial gene tree. Molecular Phylogenetics and Evolution 39 (3): 628-644 - get paper here
  • Lemos-Espinal JA, Smith GR, Rorabaugh JC 2019. A conservation checklist of the amphibians and reptiles of Sonora, Mexico, with updated species lists. ZooKeys 829: 131-160 - get paper here
  • Meyers, J.J.; Herrel, A. & Nishikawa, K. 2006. Morphological correlates of ant eating in horned lizards (Phrynosoma). Biological Journal of the Linnean Society 89: 13–24 - get paper here
  • Mosauer, Walter 1936. The reptilian fauna of sand dune areas of Vizcaino Desert and of northwestern Lower California. Occasional Papers of the Museum of Zoology, University of Michigan (329): 1-21
  • Mulcahy, Daniel G.; Allen W. Spaulding, Joseph R. Mendelson III & Edmund D. Brodie, Jr. 2006. Phylogeography of the Flat-tailed Horned Lizard (Phrynosoma mcallii) and systematics of the P. mcallii–platyrhinos mtDNA complex. Molecular Ecology 15(0): 1–20 - get paper here
  • Pianka, E. R. 1991. Phrynosoma platyrhinos. Girard: Desert Horned Lizard. Catalogue of American Amphibians and Reptiles 517: 1-4 - get paper here
  • Pianka, E. R.;Parker, W. S. 1975. Ecology of horned lizards: A review with special reference to Phrynosoma platyrhinos. Copeia 1975 (1): 141-162 - get paper here
  • Reeder,T.W. & Montanucci,R.R. 2001. Phylogenetic analysis of the horned lizards (Phrynomomatidae: Phrynosoma): evidence from mitochondrial DNA and morphology. Copeia 2001 (2): 309-323 - get paper here
  • Smith, H.M. & Taylor,E.H. 1950. An annotated checklist and key to the reptiles of Mexico exclusive of the snakes. Bull. US Natl. Mus. 199: 1-253 - get paper here
  • Stebbins,R.C. 1985. A Field Guide to Western Reptiles and Amphibians, 2nd ed. Houghton Mifflin, Boston
  • Stejneger, L.H. 1893. Annotated list of the reptiles and batrachians collected by the Death Valley Expedition in 1891, with descriptions of new species. North American Fauna, No. 7: 159-228 (+ 14 plates + 4 maps) - get paper here
  • Turner, Frederick B.;Wauer, Roland H. 1963. A survey of the herpetofauna of the Death Valley area. Great Basin Naturalist 23 (3-4): 119-128 - get paper here
  • Werning, H. 2011. Phrynosoma platyrhinos. Wüstenkrötenechse. Reptilia (Münster) 16 (90): 47-50 - get paper here
  • Werning, H. 2014. Krötenechsen – eine (sehr gute) Laune der Natur. Reptilia (Münster) 19 (107): 16-23
  • Werning, Heiko 2012. Die Reptilien und Amphibien des Südwestens. Draco 13 (50): 18-60 - get paper here
  • Woodbury, Angus Munn 1928. The reptiles of Zion National Park. Copeia 1928 (166): 14-21 - get paper here
  • Zorn, H. 2004. Die Wüste im eigenen Haus. Reptilia (Münster) 9 (48): 41-44 - get paper here
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