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Phyllodactylus lupitae CASTRO-FRANCO & URIBE-PENA, 1992

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Higher TaxaPhyllodactylidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Lane's Leaf-toed Gecko
S: Salamanquesa de Lane 
SynonymPhyllodactylus lanei lupitae CASTRO-FRANCO & URIBE-PENA 1992
Phyllodactylus lanei lupitae — DE LISLE et al. 2013
Phyllodactylus lupitae — RAMÍREZ-REYES et al. 2017
Phyllodactylus lupitae — RAMÍREZ-REYES & FLORES-VILLELA 2018
Phyllodactylus lupitae — RAMÍREZ-REYES et al. 2021 
DistributionMexico (SW Nayarit)

Type locality: Isla La Peña, Nayarit, Mexico.  
Reproductionoviparous 
TypesHolotype: EBUM 1242, adult female (UAEM), adult female 
DiagnosisDiagnosis (lupitae): This is a larger gecko along with P. lanei and P. benedettii of the P. lanei complex. It presents 28.8 tubercles from head to tail, similar to P. rupinus (28) and P. kropotkini (28.4). These counts differ from the other gecko species: P. isabelae (32.2), P. lanei (33.2), P. benedettii (27.4), P. t. magnus (37), P. muralis (33.6) and P. tuberculosus (36.4). Phyllodactylus lupitae differs clearly from the other species as it shows 25.5 third labial–snout scales, the others: P. isabelae (21.4), P. lanei (21.2), P. rupinus (20.6), P. benedettii (22.3), P. kropotkini (20.2), P. t. magnus (24.6), P. muralis (25.2) and P. tuberculosus (24.2). It is a unique species because it shows 24.6 scales across venter, the others: P. isabelae (27.8), P. lanei (30), P. rupinus (26.5), P. benedettii (29.1), P. kropotkini (30.6), P. t. magnus (27.3), P. muralis (33) and P. tuberculosus (30.2). Also presents 61.8 longitudinal scales, the others: P. isabelae (56.2), P. lanei (66), P. rupinus (63.6), P. benedettii (62.6), P. kropotkini (67.6), P. t. magnus (54.6), P. muralis (59) and P. tuberculosus (57.7). [from RAMÍREZ-REYES & FLORES-VILLELA 2018] 
CommentSubspecies: RAMÍREZ-REYES et al. 2018 diagnosed the (sub-) species of the P. lanei complex by average meristic characters (see diagnoses). This is not really useful without providing ranges (which they do provide in their Table 2, but that doesn’t have any information about sample sizes). Also, they do not provide any information about geographic variation; all specimens appear to be from small areas without any apparent sampling in between them (Fig. 1 in RAMÍREZ-REYES et al. 2018: 153). All (sub-) species appear to be genetically different, except P. lupitae and P. isabelae which are the most closely related of the P. lanei complex. The localities of lupitae and isabelae are only ~40 km apart, so closer analysis of their variation will be required to ascertain their status. 
EtymologyNamed after the wife of Ruben Castro-Franco. 
References
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