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Phyllodactylus isabelae CASTRO-FRANCO & URIBE-PENA, 1992

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Higher TaxaPhyllodactylidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Lane's Leaf-toed Gecko
S: Salamanquesa de Lane 
SynonymPhyllodactylus lanei isabelae CASTRO-FRANCO & URIBE-PENA 1992
Phyllodactylus lanei isabelae — DE LISLE et al. 2013
Phyllodactylus isabelae — RAMÍREZ-REYES et al. 2017
Phyllodactylus isabelae — RAMÍREZ-REYES & FLORES-VILLELA 2018
Phyllodactylus isabelae — RAMÍREZ-REYES et al. 2021 
DistributionMexico (Nayarit: Marietas Island)

Type locality: Marietas Island, Nayarit, Mexico.  
Reproductionoviparous 
TypesHolotype: private collection, ZUP 1152, adult female (appears to be the initials of the collector, Zeferino Uribe Peña); no other collection info provided in paper. 
DiagnosisDiagnosis (isabelae). This species clearly differs from the others because it is the smallest of the analyzed in the present study (Fig. 6). It presents 56.2 longitudinal scales on average, it differs clearly from the others: P. lupitae (61.8), P. lanei (66), P. rupinus (63.6), P. benedettii (62.6), P. kropotkini (67.6), P. t. magnus (54.6), P. muralis (59) and P. tuberculosus (57.7). Also P. isabelae presents 27.8 scales across the venter similar to P. t. magnus (27.3) and differs from the others: P. lupitae (24.6), P. lanei (30), P. rupinus (26.5), P. benedettii (29.1), P. kropotkini (30.6), P. muralis (33) and P. tuberculosus (30.2). Phyllodactylus isabelae differs clearly from the others analyzed since it has the highest number of rows of tubercles across dorsum (16.7 on average) compared to the others: P. lupitae (14.8), P. lanei (15.6), P. rupinus (14), P. benedettii (13.8), P. kropotkini (13.4), P. t. magnus (14.3), P. muralis (12.2) and P. tuberculosus (14). [from RAMÍREZ-REYES & FLORES-VILLELA 2018] 
CommentSubspecies: RAMÍREZ-REYES et al. 2018 diagnosed the (sub-) species of the P. lanei complex by average meristic characters (see diagnoses). This is not really useful without providing ranges (which they do provide in their Table 2, but that doesn’t have any information about sample sizes). Also, they do not provide any information about geographic variation; all specimens appear to be from small areas without any apparent sampling in between them (Fig. 1 in RAMÍREZ-REYES et al. 2018: 153). All (sub-) species appear to be genetically different, except P. lupitae and P. isabelae which are the most closely related of the P. lanei complex. The localities of lupitae and isabelae are only ~40 km apart, so closer analysis of their variation will be required to ascertain their status. 
EtymologyNamed after the mother of Zeferino Uribe-Peña, second author. 
References
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