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Phyllodactylus cleofasensis RAMÍREZ-REYES, BARRAZA-SOLTERO, NOLASCO-LUNA, FLORES-VILLELA & ESCO BEDO-GALVÁN, 2021

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Higher TaxaPhyllodactylidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymPhyllodactylus cleofasensis RAMÍREZ-REYES, BARRAZA-SOLTERO, NOLASCO-LUNA, FLORES-VILLELA & ESCO BEDO-GALVÁN 2021
Phyllodactylus tuberculosus — STEJNEGER 1899 (in part)
Phyllodactylus lanei — ZWEIFEL 1960 (in part)
Phyllodactylus tuberculosus saxatilis — MCDIARMID et al. 1976 (in part)
Phyllodactylus tuberculosus saxatilis — CASAS-ANDREU 1992
Phyllodactylus tuberculosus saxatilis — WOOLRICH-PIÑA et al. 2016 
DistributionMexico (Maria Cleofas Island, Nayarit)

Type locality: Mexico, Nayarit, María Cleofas Island (21.3095°N, 106.2340°W), 64 m elevation  
Reproduction 
TypesHolotype: MZFC-HE 35623, Adult female, collected by Ilse K. Barraza-Soltero and Armando H. Escobedo-Galván on on 24–25 May 2018.
Paratypes: MZFC-HE 35618-35622 and MZFC-HE 35624-35626, Six adult specimens and two juveniles. 
DiagnosisDiagnosis (and/or description): Phyllodactylus cleofasensis can be distinguished from all congeners by the following combination of characters: (1) Snout-vent length ranged from 44.8 to 77.0 mm (mean 59.5 mm), (2) has a white venter, variable dorsal coloration, and a greater number of paravertebral dorsal tubercles (mean 47.89), (3) paravertebral dorsal tubercles from head to tail (mean 47.89), (4) number of scales across the snout, starting from the 3rd labial scale (mean 26.5), and (5) number of longitudinal ventral scales from an imaginary line of the forelimbs to the cloacal opening (mean 61) (Ramírez-Reyes et al. 2021) 
CommentDiet: Orthopterans (Family Rhaphidophoridae) and coleopterans (Family Passalidae); they also consumed plant matter, arachnids, lepidopterans, scorpions (Centruroides elegans insularis), cockroaches (Blattodea: Pycnoscelus surinamensis) and remains of shed skin (Barraza-Soltero, 2019; Barraza-Soltero & Escobedo-Galván, 2020).

Similar species: P. saxatilis 
EtymologySpecific epithet is taken from the type locality María Cleofas Island, with the Latin suffix -ensis meaning, “originating from.” Specific epithet is masculine, in agreement with the gender of Phyllodactylus. 
References
  • Loc-Barragán JA, Smith GR, Woolrich-Piña GA, Lemos-Espinal JA 2024. An updated checklist of the amphibians and reptiles of Nayarit, Mexico with conservation status and comparison with adjoining States. Herpetozoa 37: 25-42 - get paper here
  • Nolasco-Luna JR, Barraza-Soltero IK, López-Montes MA, Moreno-López JA, Escobedo-Galván AH 2022. An updated checklist of the herpetofauna from Isla María Cleofas, Mexico. Check List 18(1): 241-252 - get paper here
  • Ramírez-Reyes T, Barraza-Soltero IK, Nolasco-Luna JR, Flores-Villela O, Escobedo-Galván AH 2021. A new species of leaf-toed gecko (Phyllodactylidae, Phyllodactylus) from María Cleofas Island, Nayarit, Mexico. ZooKeys 1024: 117-136 - get paper here
 
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