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Phyllodactylus nolascoensis DIXON, 1964

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Higher TaxaPhyllodactylidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Nolasco Leaf-toed Gecko
S: Salamanquesa de Nolasco 
SynonymPhyllodactylus homolepidurus nolascoensis DIXON 1964: 42
Phyllodactylus homolepidurus nolascoensis — WERMUTH 1965: 137
Phyllodactylus homolepidurus nolascoensis — DE LISLE et al. 2013
Phyllodactylus nolascoensis — LEMOS-ESPINAL et al. 2019
Phyllodactylus nolascoensis — RAMÍREZ-REYES et al. 2021 
DistributionMexico (San Pedro Nolasco Island)

Type locality: San Pedro Nolasco Island, 15 km WSW of Punta San Pedro, Sonora, Mexico.  
Reproductionoviparous 
TypesHolotype: CAS; Paratypes SDNHM = SDSNH 6840, 6841 
DiagnosisDIAGNOSIS: An average-sized gecko, snout-vent length 45 to 62 (53,0) mm" with 14 to 16 longitudinal rows of enlarged dorsal tubercles. which are more numerous than those found in the nominal subspecies, Number of paravertebral tubercles 41 to 48 rather than 31 to 41, as in P. homolepidurus homolepidurus (Dixon 1964).

DESCRIPTION OF HOLOTYPE: Rostral slightly more than twice as wide as high, its upper edge shaped as a broad "W" with a distinct median groove one-half the depth of rostral; internasals about as wide as longo roughly rectangular» point of angle extending posteriorly; internasals bordered posteriorly by 5 granules and postnasal of each side; nostril surrounded by internasal» 2 postnasals, rostral, and first supralabial; supralabial in moderate contact with ventral edge of nostril; deep depression in frontal region; 17 scales across snout between second labial, 21 between third labials; 14 scales between eye and nostril; second supralabial separated from nostril by 2 granules and postnasal; diameter of eye contained in snout length slightly less than 2 times; 17 scales between anterior edge of orbitsD 19 interorbital scales; 3 rows of granules and one larger outer row of scales bordering eyelid. last 2 or 3 scales of latter row pointed, spine-like; ear opening not denticulateo scales bordering anterior edge about twice the size of granules bordering posterior edge; diameter of ear contained in diameter of eye about two and one-half times; 21 rows of granules between posterior border of eye and anterior edge of ear; posterodorsal surface of head with small granules» slightly smaller than orbital scales, and intermixed with scatteredo larger rounded tubercles; 7 supralabials and 6 infralabials to point below center of eye; mental bell-shaped» slightly wider than long, followed by 2 postmentals which are twice as long as wideD contacting only first labial of each side; 10 scales bordering postmentals posteriorly» followed by second transverse row of 12 smaller scales; first and second infralabials of equal size. Upper arm with flat. imbricate scales on dorsal surface» posteroventral surface granular; posteroventral surface of forearm with rounded granules; scaes of the dorsal surface rounded and imbricate, with 1 or 2 intermixed tubercles; scales on dorsal, anteodorsal, and ventral surface of thigh circular, posteroventral surface granular with 1 to 3 tubercles intermixed; dorsal surface of lower leg with small conical scales with several larger tubercles intermixed; terminal pads of toes and fingers slightly longer than wide, truncate; claws completely hidden between terminal pads when viewed from side and from below; lamellae formula for hand 8-9-10-11-9, foot 7-9-13-14-11 0 Dorsum with 14 longitudinal rows of enlarged trihedral tubercles, 10 of which reach to rear of head, 6 reach base of tail; 44 paravertebral tubercles, 24 between axilla and groin; median two longitudinal rows of enlarged dorsal tubercles separated from each other by 5 to 6 rows of granules; each tubercle separated from preceding tubercle by 1 to 3 granules; 5 rows of granules separate ventral scales from first row of tubercles; 30 scales across venter, 59 from gular region to anus; scales of venter abruptly differentiated from lateral granules; 4 postanal tubercles on one side, 3 on the other, all distinct and moderately elevated; postanal region swollen; tail regenerated, without tubercles (Dixon 1964). 
CommentGrismer (1999) synonymized Phyllodactylus homolepidurus nolascoensis with Phyllodactylus homolepidurus without giving specific reasons. 
EtymologyNamed after the type locality. 
References
  • Dixon, J. R. 1964. The systematics and distribution of lizards of the genus Phyllodactylus in North and Central America. New Mexico State Univ. Sci. Bull. 64: 1-139 - get paper here
  • Enderson, Erik F.; Thomas R. Van Devender, Robert L. Bezy 2014. Amphibians and reptiles of Yécora, Sonora and the Madrean Tropical Zone of the Sierra Madre Occidental in northwestern Mexico. Check List 10 (4): 913-926 - get paper here
  • Goldberg, Stephen R. 2008. Note on Reproduction of the Sonoran Leaf-toed Gecko, Phyllodactylus homolepidurus (Squamata: Gekkonidae) from Sonora, Mexico. Bull. Maryland Herp. Soc. 44 (2): 46-48 - get paper here
  • Grismer, L. Lee. 1999. An evolutionary classification of reptiles on islands in the Gulf of California, México. Herpetologica 55 (4): 446-469 - get paper here
  • Hardy, L.M., & McDiarmid, R.W. 1969. The amphibians and reptiles of Sinaloa, Mexico. Univ. Kansas Publ. Mus. Nat. Hist. 18 (3): 39-252. - get paper here
  • Jones, L.L. & Lovich, R.E. 2009. Lizards of the American Southwest. A photographic field guide. Rio Nuevo Publishers, Tucson, AZ, 568 pp. [review in Reptilia 86: 84] - get paper here
  • Lemos-Espinal JA, Smith GR 2020. A checklist of the amphibians and reptiles of Sinaloa, Mexico with a conservation status summary and comparisons with neighboring states. ZooKeys 931: 85-114 - get paper here
  • Lemos-Espinal JA, Smith GR, Rorabaugh JC 2019. A conservation checklist of the amphibians and reptiles of Sonora, Mexico, with updated species lists. ZooKeys 829: 131-160 - get paper here
  • Nevárez-de los Reyes; Manuel, David Lazcano, Javier Banda-Leal and Ian Recchio 2014. Notes on Mexican Herpetofauna 22: Herpetofauna of the Continental Portion of the Municipality of Hermosillo, Sonora, Mexico. Bull. Chicago Herp. Soc. 49(8):105-115 - get paper here
  • Ramírez-Reyes T, Barraza-Soltero IK, Nolasco-Luna JR, Flores-Villela O, Escobedo-Galván AH 2021. A new species of leaf-toed gecko (Phyllodactylidae, Phyllodactylus) from María Cleofas Island, Nayarit, Mexico. ZooKeys 1024: 117-136 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Smith, H.M. 1935. Miscellaneous notes on Mexican lizards. Univ. Kansas Sci. Bull. 22: 119-156 - get paper here
  • Smith, H.M. & Taylor,E.H. 1950. An annotated checklist and key to the reptiles of Mexico exclusive of the snakes. Bull. US Natl. Mus. 199: 1-253 - get paper here
  • Taylor, Edward Harrison 1938. Notes on the herpetological fauna of the Mexican state of Sonora. Univ. Kansas Sci. Bull. 24 (19): 475-503 [1936] - get paper here
  • Zweifel,R.G. and Norris,K.S. 1955. Contributions to the herpetology of Sonora, Mexico:Descriptions of new subspecies of snakes (Micruroides euryxanthus and Lampropeltis getulus) and miscellaneous collecting notes. American Midland Naturalist 54: 230-249 - get paper here
 
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