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Phymaturus aguanegra LOBO, LASPIUR & ACOSTA, 2013

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Higher TaxaLiolaemidae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common Names 
SynonymPhymaturus aguanegra LOBO, LASPIUR & ACOSTA 2013 
DistributionNW Argentina

Type locality: Paso Agua Negra, Iglesia department, San Juan province, Argentina. 30°23’ S; 69°34’ W. 2900 m elevation.  
Reproductionviviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: MCN-UNSa (= MCN) 975. Male. April 2007. A. Laspiur, E. Sanabria & L. Quiroga collectors. Paratypes: MCN (969, 973-975, 977, 979, 982, 984, 988, 990-991, 995 and 971-972, 976, 978, 980, 981, 983, 986-987, 989, 992-993). 13 males and 14 females. Same data as holotype. 
DiagnosisDiagnosis: Phymaturus aguanegra sp. nov. differs from all other members (P. antofagastensis, P. denotatus, P. laurenti, P. punae, P. extrilidus, P. mallimaccii and P. paihuanense) exhibiting the following combination of diagnostic characters: a complete melanism over the dorsum of neck, the presence of enlarged scales at the base of tail in males, having strongly keeled tarsal scales, lacking enlarged scales on the anterior margin of the antehumeral fold and centre of chest, females without flank coloration, a vertebral dark gray stripe usually present on the dorsum, females exhibiting a tricolor dorsal pattern, with two types of brown and scattered ferriferous oxide spots, and the absence of a scapular spot.


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CommentAbundance: only known from its original description (Meiri et al. 2017). 
EtymologyThe specific epithet is given after its type locality. [“Agua Negra”] toponym, which refers to the dark water aspect of Andean streams near the type locality. 
References
  • LASPIUR, A., MEDINA, M., AUSAS, M. S., ACOSTA, J. C., KRENZ, J. D., & IBARGÜENGOYTÍA, N. R. 2024. Thermal niches and activity periods in syntopic Phymaturus and Liolaemus lizards from the Andes, Argentina. Anais da Academia Brasileira de Ciências, 96, e20191190 - get paper here
  • Lobo, F., Barrasso, D. A., Hibbard, T. and Basso, N. G. 2015. On the evolution and diversification of an Andean clade of reptiles: combining morphology and DNA sequences of the palluma group (Liolaemidae: Phymaturus). Zoological Journal of the Linnean Society 176:648–673, doi: 10.1111/zoj.12335 [2016]
  • LOBO, FERNANDO; ALEJANDRO LASPIUR, JUAN CARLOS ACOSTA 2013. Description of new andean species of the genus Phymaturus (Iguania: Liolaemidae) from Northwestern Argentina. Zootaxa 3683: 117–132 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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