Phymaturus aguanegra LOBO, LASPIUR & ACOSTA, 2013

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Higher Taxa	Liolaemidae, Iguania, Sauria, Squamata (lizards)
Subspecies
Common Names
Synonym	Phymaturus aguanegra LOBO, LASPIUR & ACOSTA 2013
Distribution	NW Argentina Type locality: Paso Agua Negra, Iglesia department, San Juan province, Argentina. 30°23’ S; 69°34’ W. 2900 m elevation.
Reproduction	viviparous (not imputed, fide Zimin et al. 2022)
Types	Holotype: MCN-UNSa (= MCN) 975. Male. April 2007. A. Laspiur, E. Sanabria & L. Quiroga collectors. Paratypes: MCN (969, 973-975, 977, 979, 982, 984, 988, 990-991, 995 and 971-972, 976, 978, 980, 981, 983, 986-987, 989, 992-993). 13 males and 14 females. Same data as holotype.
Diagnosis	Diagnosis: Phymaturus aguanegra sp. nov. differs from all other members (P. antofagastensis, P. denotatus, P. laurenti, P. punae, P. extrilidus, P. mallimaccii and P. paihuanense) exhibiting the following combination of diagnostic characters: a complete melanism over the dorsum of neck, the presence of enlarged scales at the base of tail in males, having strongly keeled tarsal scales, lacking enlarged scales on the anterior margin of the antehumeral fold and centre of chest, females without flank coloration, a vertebral dark gray stripe usually present on the dorsum, females exhibiting a tricolor dorsal pattern, with two types of brown and scattered ferriferous oxide spots, and the absence of a scapular spot. Additional details (3463 characters) are available for collaborators and contributors. Please contact us for details.
Comment	Abundance: only known from its original description (Meiri et al. 2017).
Etymology	The specific epithet is given after its type locality. [“Agua Negra”] toponym, which refers to the dark water aspect of Andean streams near the type locality.
References	LASPIUR, A., MEDINA, M., AUSAS, M. S., ACOSTA, J. C., KRENZ, J. D., & IBARGÜENGOYTÍA, N. R. 2024. Thermal niches and activity periods in syntopic Phymaturus and Liolaemus lizards from the Andes, Argentina. Anais da Academia Brasileira de Ciências, 96, e20191190 - get paper here Lobo, F., Barrasso, D. A., Hibbard, T. and Basso, N. G. 2015. On the evolution and diversification of an Andean clade of reptiles: combining morphology and DNA sequences of the palluma group (Liolaemidae: Phymaturus). Zoological Journal of the Linnean Society 176:648–673, doi: 10.1111/zoj.12335 [2016] LOBO, FERNANDO; ALEJANDRO LASPIUR, JUAN CARLOS ACOSTA 2013. Description of new andean species of the genus Phymaturus (Iguania: Liolaemidae) from Northwestern Argentina. Zootaxa 3683: 117–132 - get paper here Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
External links	IUCN Red List Google images

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