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Pinoyscincus coxi (TAYLOR, 1915)

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Higher TaxaScincidae, Sphenomorphinae (Sphenomorphini), Scincoidea, Sauria, Squamata (lizards)
SubspeciesPinoyscincus coxi coxi (TAYLOR 1915)
Pinoyscincus coxi divergens (TAYLOR 1922) 
Common NamesE: Cox's Sphenomorphus 
SynonymSphenomorphus coxi coxi TAYLOR 1915: 100
Sphenomorphus coxi TAYLOR 1915: 100
Lygosoma (Hinulia) jagori PETERS 1864: 54 (part.)
Lygosoma (Sphenomorphus) coxi — BROWN & ALCALA 1970: 113
Sphenomorphus coxi coxi — BROWN & ALCALA 1980: 203
Pinoyscincus coxi — LINKEM, DIESMOS & BROWN 2011

Pinoyscincus coxi divergens TAYLOR 1922
Sphenomorphus jagori divergens TAYLOR 1922: 194
Lygosoma (Sphenomorphus) jagori — SMITH 1937: 220 (part.)
Sphenomorphus coxi divergens — BROWN & ALCALA 1980: 206
Sphenomorphus coxi divergens — BROWN et al. 1995
Sphenomorphus coxi divergens — LAGAT 2009
Pinoyscincus coxi divergens (by implication)
Pinoyscincus coxi divergens — MENESES et al. 2022 
DistributionPhilippines (coxi: Mindanao, Leyte, Samar, Camiguin; divergens: SW Luzon, Marinduque, Mindoro)

divergens: widely distributed in the Philippines (incl. Luzon, Marinduque, Mindoro (Crombie, 1994). Also recorded from Mindoro, Luzon, Polillo, Leyte, Samar, Camiguin, Mindanao, Panay, and Sulu Islands).

Type locality: Bunawan (Bunauan), Agusan Province, Mindanao Island [coxi].
Type locality: Mt. Makiling, Luzon, and Naujan and Calapan, Mindoro Island [divergens].  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: lost, BSMP 1782, collected E.H. Taylor, vi.1912; lost fide BROWN & ALCALA 1980: 204
Syntypes: CM P1758-66, Sumagui & Pocamil, Mindoro, collected E.H. Taylor v.1916; P1767-78, Calapan & San Isidro, Mindoro, collected E.H. Taylor, xi.1916. UMMZ 65965, Calapan, Mondoro, collected E.H. Taylor, x.1916, MCZ 20134, Sumagui & Pocamil, Mindoro, collected E.H.Taylor v.1915, and MCZ 20135, Calapan & San Isidro, Mindoro, collected E.H. Taylor, xi.1916 are also possibly syntypes, although Taylor (1922) only listed 9 specimens from Sumagui and 12 from Calapan. Whereabouts of the remaining 10 syntypes (4 from Mt Maquiling, Luzon; 6 from Lake Naujan, Mindoro) unknown, although Marx (1972) refers to 4 "cotypes" of unspecified locality in FMNH [divergens] 
DiagnosisDiagnosis:-Frontoparietal single; prefrontals not in contact; 5 supraoculars; adpressed hind legs fail to reach beyond the elbow; frontal as broad as, or slightly broader than, the supraocular region; last 3 labials with distinct yellow spots.” (Taylor 1915: 100)


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Comment 
EtymologyNamed after Dr. Alvin J. Cox, former Director of the Philippines Bureau of Science, Manila, ca. 1910. 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Beukema, W. 2011. Herpetofauna of disturbed forest fragments on the lower Mt. Kitanglad Rnage, Mindanao Isand, Philippines. Salamandra 47 (2): 90-98 - get paper here
  • Brown, R.M. et al. 2010. Species boundaries in Philippine montane forest skinks (Genus Sphenomorphus): three new species from the mountains of Luzon and clarification of the status of the poorly known S. beyeri, S. knollmanae, and S. laterimaculatus. Scient. Pap. Nat. Hist. Mus. Univ. Kansas (42): 1-27
  • Brown, WC. & A.C. ALCALA 1980. Philippine Lizards of the family Scincidae. Silliman Univ. Nat. Sci., Dumaguete City, Mon., Ser. 2: i-xi + 1-246.
  • Brown,R.M. & Ferner,J.W. & Sison,R.V. 1995. Rediscovery and redescription of Sphenomorphus beyeri Taylor (Reptilia: Lacertilia: Scincidae) from the Zambales Mountains of Luzon, Philippines. Proc. Biol. Soc. Washington 108 (1): 6-17 - get paper here
  • Decena, S. C. P., Macasait Jr, D. R., & Arguelles, M. S. 2023. Species Accounts, Assemblage, and Microhabitats of Amphibians and Reptiles of Northeastern Leyte, Philippines. Philippine Journal of Science, 152(1), 1-34
  • Ferner, John W., Rafe M. Brown, Rogelio V. Sison and Robert S. Kennedy 2000. The amphibians and reptiles of Panay Island, Philippines. Asiatic Herpetological Research 9: 1-37 - get paper here
  • Gaulke, M. 2011. The herpetofauna of Panay Island, Philippines. Edition Chimaira, 390 pp.
  • Lagat, R.D. 2009. A TAXONOMIC ACCOUNT OF LIZARDS ALONG ESTABLISHED TRAILS IN MTS. PALAY-PALAY MATAAS-NA-GULOD PROTECTED LANDSCAPE, LUZON ISLAND, PHILIPPINES. Philippine Journal of Systematic Biology 3: 17-28 - get paper here
  • Linkem, Charles W.; Arvin C. Diesmos, Rafe M. Brown 2011. Molecular systematics of the Philippine forest skinks (Squamata: Scincidae: Sphenomorphus): testing morphological hypotheses of interspecific relationships. Zoological Journal of the Linnean Society 163: 1217–1243 - get paper here
  • Linkem, Charles W.;Kyle Miller Hesed, Arvin C. Diesmos, Rafe M. Brown 2010. Species boundaries and cryptic lineage diversity in a Philippine forest skink complex (Reptilia; Squamata; Scincidae: Lygosominae). Molecular Phylogenetics and Evolution 56 (2): 572-585 - get paper here
  • Meneses CG, Siler CD, Alviola PA, Gonzalez JCT, Balatibat JB, Natividad CA, Brown RM 2022. Amphibian and reptile diversity along a ridge-to-reef elevational gradient on a small isolated oceanic island of the central Philippines. Check List 18(5): 941-948 - get paper here
  • Mohagan, Alma B.; Olga M. Nuñeza, Angel C. Alcala, Jose A. Escarlos Jr., Arturo G. Gracia Jr., Eric Claire T. Selpa, Leonil Jun B. Baguhin, Fulgent P. Coritico & Victor B. Amoroso 2019. Species richness and endemism of Reptilian Fauna in four Long-Term Ecological Research sites in Mindanao, Philippines. Biodiversity Journal, 2019, 10 (3): 237–248 - get paper here
  • Peters,W.C.H. 1864. Die Eidechsenfamilie der Scincoiden, insbesondere über die Schneider'schen, Wiegmann'schen und neue Arten des zoologischen Museums. Monatsber. K. Akad. Wiss. Berlin, 1864: 44-58 - get paper here
  • Rosales, A., Suhian, J. P., Gamalinda, E., Paz, S., & Solania-Naling, C. 2023. Herpetofauna of forest fragments within the Agata Mining Ventures Incorporated (AMVI), Agusan del Norte, Caraga Region, Philippines. Journal of Ecosystem Science and Eco-Governance, 5(2), 54-65 - get paper here
  • Sanguila MB, Cobb KA, Siler CD, Diesmos AC, Alcala AC, Brown RM 2016. The amphibians and reptiles of Mindanao Island, southern Philippines, II: the herpetofauna of northeast Mindanao and adjacent islands. ZooKeys 624: 1–132, doi: 10.3897/zookeys.624.9814 - get paper here
  • Smith, M.A. 1937. A review of the genus Lygosoma (Scincidae: Reptilia) and its allies. Records of the Indian Museum 39 (3): 213-234
  • Taylor, E.H. 1915. New species of Philippine lizards. Philippine Journal of Science. 10: 89-109 - get paper here
  • Taylor, E.H. 1922. The lizards of the Philippine Islands. Department of Agriculture and Natural Resources, Bureau of Science, Government of the Philippine Islands, Manila, Publication no. 17: 269 pp. - get paper here
  • VENTURINA, Russell Evan L.;Ynigo Luis C. DEL PRADO,Rhim Ayyah C. KAMIR, Marlon N. BALMORES and Arvin C. DIESMOS 2020. A Revised Checklist of Amphibians and Reptiles in Camiguin Sur, Misamis Oriental, Mindaanao, Philippines. Asian Herpetological Research 11 (1): 28-43 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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