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Plica umbra (LINNAEUS, 1758)

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Higher TaxaTropiduridae, Iguania, Sauria, Squamata (lizards)
SubspeciesPlica umbra umbra (LINNAEUS 1758)
Plica umbra ochrocollaris (SPIX 1825) 
Common NamesE: Blue-Lipped Tree Lizard, Harlequin Racerunner
Portuguese: Calango, Calango-da-Árvore, Iguaninha, Lagarto-de-Árvore, Papa-Vento, Tamaquaré 
SynonymLacerta Umbra LINNAEUS 1758: 207
Iguana umbra — MEYER 1795: 17
Iguana umbra — LATREILLE 1801
Agama umbra — DAUDIN 1802
Ophryessa ochrocollaris — BOIE 1825
Ophryessa umbra — FITZINGER 1826
Uraniscodon (Pneustes) umbra — KAUP 1827
Hypsibatus umbra — WAGLER 1830
Ophyessa [sic] (Plica) picta — GRAY 1831
Hypselophus umbra — WIEGMANN 1835
Uperanodon ochrocollare — DUMÉRIL & BIBRON 1837
Hypsibatus (Hypsibatus) umbra — FITZINGER 1843
Uraniscodon umbra — GRAY 1845
Uperanodon Ochrocollare — DUMÉRIL, BIBRON & DUMÉRIL 1854: 267
Plica umbra — O’SHAUGHNESSY 1881: 245
Uraniscodon umbra — BOULENGER 1885: 179
Tropidurus unicarinatus WERNER 1899: 480
Plica umbra — BURT & BURT 1933: 40
Plica umbra — PETERS et al. 1970: 230
Tropidurus umbra — FROST 1992
Tropidurus umbra — DIRKSEN & DE LA RIVA 1999
Plica umbra umbra — GORZULA & SEÑARIS 1999
Tropidurus umbra — HARVEY & GUTBERLET 2000
Plica umbra — FROST et al. 2001
Tropidurus umbra — LEHR 2002: 88
Tropidurus umbra — WHITHWORTH & BEIRNE 2011
Plica umbra — MURPHY & JOWERS 2013

Plica umbra ochrocollaris (SPIX 1825)
Lophyrus ochrocollaris SPIX 1825: 10
Uperanodon ochrocollare — DUMÉRIL & BIBRON 1837: 248
Hyperanodon ochrocollaris — COPE 1876: 170
Hyperanodon peltigerus COPE 1875: 170
Tropidurus holotropis BOULENGER 1912: 420
Uraniscodon tuberculatum ANDERSSON 1918: 2
Plica tuberculatum ANDERSSON 1918
Plica umbra ochrocollaris — PETERS et al. 1970: 231
Plica umbra ochrocollaris — DIXON & SOINI 1975
Plica umbra ochrocollaris — DUELLMAN 1978: 206
Plica umbra ochrocollaris — PRUDENTE et al. 2013
Plica umbra ochrocollaris — OLIVEIRA et al. 2016
Plica umbra ochrocollaris — RIBEIRO-JUNIOR & AMARAL 2016 
DistributionColombia, Venezuela, Guyana, Suriname, French Guiana,
Brazil (Pará, Acre, Amazonas, Rondônia etc.), Bolivia (Beni, La Paz, Pando, Santa Cruz), N Peru (Pasco etc.), Ecuador

ochrocollaris: Bolivia, Brazil (Mato Grosso, Amazonas, Pará)

Type locality: “in Meridionalibus”  
Reproductionoviparous 
TypesSyntypes: NRM (NHRM) 111 (2 specimens), UUZM 73
Lectotype: RMNH 2899, designated by Hoogmoed & Gruber (1983). Original syntypes: ZSM 747/1920 (lost), sex unknown, from "in sylvis fluminis Amazonum", collected by Spix and Martius expedition to Brazil 1817-1820 [ochrocollaris]
Holotype: ANSP 13095 [Hyperanodon peltigerus]
Syntypes: NMW [Tropidurus unicarinatus] 
DiagnosisDiagnosis: As generic diagnosis, and in addition the following features: body roughly cylindrical, no tufts of spinose scales on neck, scales around midbody 43-69. Maximum SVL c. 100 mm. Mottled green and brown, one or the other colour predominating; transverse dark, spotted bands may be present (Avila-Pires 1995: 172).


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CommentSynonymy partly after Peters et al. 1970.

Plica umbra ochrocollaris is comprised of five lineages (Oliveira et al. 2016).

Diet: at least some populations of this species live almost exclusively from ants (Lucas et al. 2023). 
EtymologyNamed after Latin umbra, a shade, shadow. (LINNAEUS 1758, Esteban Lavilla, pers. comm., April 2024) 
References
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