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Podarcis milensis (BEDRIAGA, 1882)

IUCN Red List - Podarcis milensis - Vulnerable, VU

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Higher TaxaLacertidae, Lacertinae, Lacertini, Sauria, Lacertoidea, Squamata (lizards)
SubspeciesPodarcis milensis adolfjordansi (BUCHHOLZ 1962)
Podarcis milensis gerakuniae (MÜLLER 1938)
Podarcis milensis milensis (BEDRIAGA 1882) 
Common NamesE: Milos Wall Lizard
G: Milos-Mauereidechse 
SynonymLacerta muralis fusca var. milensis BEDRIAGA 1882: 98
Lacerta muralis fusca var. nigrogularis WERNER 1899: 835
Lacerta erhardii schweizeri MERTENS 1934
Lacerta muralis milensis — WERNER 1935: 92
Podarcis milensis — BEUTLER & HECKES 1986
Podarcis milensis schweizeri — ENGELMANN et al 1993
Podarcis milensis — ENGELMANN et al 1993
Podarcis milensis — POULAKAKIS et al. 2005
Podarcis milensis — PSONIS et al. 2021

Podarcis milensis adolfjordansi (BUCHHOLZ 1962)
Lacerta muralis adolfjordansi BUCHHOLZ 1962: 218
Podarcis milensis adolfjordansi — SINDACO & JEREMČENKO 2008

Podarcis milensis gerakuniae (MÜLLER 1938)
Podarcis milensis gerakuniae — SINDACO & JEREMČENKO 2008

Podarcis milensis milensis (BEDRIAGA 1882)
Podarcis milensis milensis — SINDACO & JEREMČENKO 2008 
DistributionGreece (Milos: Kimolos, western Cyclades, Ananes Archipelago, Islands Falkonera and Velopoula)

adolfjordansi: Ananes Island (W of Milos)

gerakuniae: Gerakunia (= Falconera) Island; Type locality: gerakuniae] “Insel Gerakúnia, nordwestl. von Erimomilos, Kykladen”

nigrogularis: Milos  
TypesTypes: BMNH 1946.9.1.59-60 (and possibly additional specimens).
Holotype: lost, was ZSM (SLM) 2409, male, collected by H. Schweizer, 05.1935. Apparently the whole type series is lost at ZSM fide Franzen & Glaw 2007 [gerakuniae]
Holotype: ZFMK 1854, paratypes in ZFMK and ZMB [adolfjordansi] 
CommentSynonymy: BEUTLER & HECKES 1986 list most of the names in the synonymy under P. milensis without saying explicitly they are synonyms or valid subspecies. We only recognized those listed by SINDACO & JEREMČENKO 2008. The paper by Bedriaga 1882 is not explicitely mentioned in the Zoological Record of 1882. 
EtymologyNamed after the type locality (Milos). 
  • Adamopoulou, C. & E.D. Valakos 2000. Small clutch size in a Mediterranean endemic Lacertid (Podarcis milensis) Copeia 2000 (2): 610-614. - get paper here
  • Andrea Villa, Massimo Delfino 2019. A comparative atlas of the skull osteology of European lizards (Reptilia: Squamata). Zoological Journal of the Linnean Society 187 (3): 829–928 - get paper here
  • Bedriaga, J. von. 1882. Die Amphibien und Reptilien Griechenlands. Bulletin de la Société Impériale des Naturalistes de Moscou, 56 (2): 242-310; (3) 43-103, (4) 278-344 [published in 3 parts] - get paper here
  • Beutler,A. & Heckes,U. 1986. Podarcis milensis - Milos-Mauereidechse. In: Böhme,W. (ed.), Handbuch der Reptilien und Amphibien Europas, Band 2/II., Echsen III (Podarcis). Aula-Verlag Wiesbaden, pp. 142-154
  • Böhme, Wolfgang 2014. Herpetology in Bonn. Mertensiella 21. vi + 256 pp. - get paper here
  • Broggi, Mario F. 2014. The herpetofauna of Kimolos (Milos Archipelago, Greece). Herpetozoa 27 (1/2): 102-103 - get paper here
  • Buchholz, K.F. 1962. Die Mauereidechse von Ananes, Milos-Archipel. Bonner zoologische Beiträge 13 (1/3): 216-218 - get paper here
  • Engelmann, W.E. et al. 1993. Lurche und Kriechtiere Europas. Neumann Verlag (Radebeul, Germany), 440 pp.
  • Kwet, A. 2010. Reptilien und Amphibien Europas. Kosmos-Verlag, Stuttgart, 252 pp.
  • Kwet, Axel & Benny Trapp 2014. Liste der Reptilien Europas. Draco 15 (60): 72-79 - get paper here
  • Müller, L. 1938. Über die Mauereidechse der Insel Gerakunia. Blätter für Aquarien- und Terrarienkunde 1938 (3): 38-40 - get paper here
  • Poulakakis, N., P. Lymberakis; E. Valakos; E. Zouros and M. Mylonas 2005. Phylogenetic relationships and biogeography of Podarcis species from the Balkan Peninsula, by bayesian and maximum likelihood analyses of mitochondrial DNA sequences. Molecular Phylogenetics and Evolution 37 (3): 845-857 - get paper here
  • Poulakakis, N.; P. Lymberakis, A. Antoniou, D. Chalkia, E. Zouros, M. Mylonas and E. Valakos 2003. Molecular phylogeny and biogeography of the wall-lizard Podarcis erhardii (Squamata: Lacertidae). Molecular Phylogenetics and Evolution 28 (1): 38-46 - get paper here
  • POULAKAKIS, N.; P. LYMBERAKIS; E. VALAKOS; P. PAFILIS; E. ZOUROS and M. MYLONAS 2005. Phylogeography of Balkan wall lizard (Podarcis taurica) and its relatives inferred from mitochondrial DNA sequences. Molecular Ecology 14: 2433–2443 - get paper here
  • Psonis, N., Antoniou, A., Karameta, E., Darriba, D., Stamatakis, A., Lymberakis, P., & Poulakakis, N. 2021. The wall lizards of the Balkan peninsula: tackling questions at the interface of phylogenomics and population genomics. Molecular Phylogenetics and Evolution, 107121 - get paper here
  • Psonis, N., Antoniou, A., Kukushkin, O., Jablonski, D., Petrov, B., Crnobrnja – Isailović, J., Sotiropoulos, K., Gherghel, I., Lymberakis, P., Poulakakis, N. 2016. Hidden diversity in the Podarcis tauricus (Sauria, Lacertidae) species subgroup in the light of multilocus phylogeny and species delimitation. Molecular Phylogenetics and Evolution 106: 6-17 [print version published 2017] - get paper here
  • Schmid, A. 2015. Feta-Käse, Sonne und Schlangen. Willkommen auf Milos! Reptilia (Münster) 20 (113): 90-95 - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Trutnau, L. 1975. Europaische Amphibien und Reptilien. Belser Verlag, Stuttgart 212 pp.
  • Valakos, Efstratios D.; Andriani Kourkouli, Margarita Skopeliti, Panayiotis Pafilis, Nikolaos Poulakakis, Ioannis F. Voutsas, Petros Lymberakis, Chrisi Simou, Wolfgang Voelter and Ourania E. Tsitsilonis 2007. Combining immunological and molecular data to assess phylogenetic relations of some Greek Podarcis species. Comparative Biochemistry and Physiology Part B: Biochemistry and Molecular Biology 147 (1): 1-10 - get paper here
  • Werner, F. 1899. Beiträge zur Kenntnis der Reptilien- und Batrachierfauna der Balkanhabinsel. Wiss. Mitt. Bosn. Herceg. 6: 828-837 - get paper here
  • Werner,F. 1935. Reptilien der Ägäischen Inseln. Sitzungsber. Akad. Wiss. Wien 144: 81-117 - get paper here
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