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Proctoporus bolivianus WERNER, 1910

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Higher TaxaGymnophthalmidae (Cercosaurinae), Sauria, Gymnophthalmoidea, Squamata (lizards)
Subspecies 
Common NamesE: Bolivian Lightbulb Lizard 
SynonymProctoporus bolivianus WERNER 1910: 30
Proctoporus longicaudatus ANDERSSON 1914
Proctoporus bolivianus — DIRKSEN & DE LA RIVA 1999
Proctoporus bolivianus — DOAN & SCHARGEL 2003
Proctoporus subsolanus DOAN, CASTOE & ARIZABAL ARRIAGA 2005
Proctoporus bolivianus — GOICOECHEA et al. 2012 
DistributionN Bolivia (La Paz), S Peru (Puno), elevation 2100–3743 m

Type locality: Sorata, Bolivia; Neotype locality: Millipaya, 12 km S of Sorata (15°51′30.2′′S/68°37′37.6′′W), Province Larecaja, Department of La Paz, Bolivia, 3743 m elevation.

subsolanus: Peru (Puno); Type locality: from the town of Sandia (14.34275° S, 69.46274° W), Province of Sandia, Department of Puno, Peru; 2100 m elevation.  
Reproductionoviparous 
TypesNeotype: CBF 3437 (fide Goicoechea et al. 2013); Holotype: ZMH (apparently lost)
Holotype: UTA R-52944; NCBI tax ID 321038 [subsolanus] 
DiagnosisDiagnosis: in GOICOECHEA et al. 2013.


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CommentSynonymy after PETERS & DONOSO-BARROS 1970 and GOICOECHEA et al. 2013. Oreosaurus lacertus STEJNEGER 1913: 546 has been revalidated by GOICOECHEA et al. 2013.

Habitat: during the day under logs and stones, occupying disturbed areas and agricultural fields. 
EtymologyProctoporus bolivianus has been named after its range. P. subsolanus has been named after the Latin word for “eastern”, referring to the distribution of the species in the easternmost flank of the Andes Mountains in S Peru. 
References
  • Andersson, L.G. 1914. A new Telmatobius and new teiid lizards from South America. Arkiv för Zoologi 9 (3): 1-12 - get paper here
  • Barbour, Thomas & Noble, G. K. 1921. Amphibians and reptiles from southern Peru collected by the Peruvian expedition of 1914-1915 under the auspices of Yale University and the National Geographic Society. Proc. US Natl. Mus. 58 (2352): 609-620 [1920] - get paper here
  • CATENAZZI, A., LEHR, E. & VON MAY, R. 2013. The amphibians and reptiles of Manu National Park and its buffer zone, Amazon basin and eastern slopes of the Andes, Peru. Biota Neotropica 13 (4): 269-283
  • Dirksen, L. & De la Riva, I. 1999. The lizards and amphisbaenians of Bolivia (Reptilia, Squamata): checklist, localities, and bibliography. Graellsia 55: 199-215 - get paper here
  • Doan, T. M. & Castoe, T.A. 2003. Using morphological and molecular evidence to infer species boundaries within Proctoporus bolivianus Werner (Squamata: Gymnopthalmidae). Herpetologica 59 (3): 432–449 - get paper here
  • Doan, T.F.; Castoe, T.A. & Arizábal Arriaga, W. 2005. Phylogenetic relationships of the genus Proctoporus sensu stricto (Squamata: Gymnophthalmidae), with a new species from Puno, Southeastern Peru. Herpetologica 61 (3): 325-336 - get paper here
  • Doan, T.M. 2003. A south-to-north biogeographic hypothesis for Andean speciation: evidence from the lizard genus Proctoporus (Reptilia, Gymnophthalmidae). Journal of Biogeography 30: 361–374 - get paper here
  • Doan, T.M. & Schargel, W.E. 2003. Bridging the gap in Proctoporus distribution: a new species (Squamata: Gymnophthalmidae) from the Andes of Venezuela. Herpetologica 59 (1): 68-75 - get paper here
  • Goicoechea, Noemí; José M. Padial, Juan C. Chaparro, Santiago Castroviejo-Fisher, Ignacio De la Riva 2012. Molecular phylogenetics, species diversity, and biogeography of the Andean lizards of the genus Proctoporus (Squamata: Gymnophthalmidae). Molecular Phylogenetics and Evolution, 65 (3): 953–964 [print version published in 2013], http://dx.doi.org/10.1016/j.ympev.2012.08.017
  • Köhler, G. and E. Lehr 2004. Comments on Euspondylus and Proctoporus (Squamata: Gymnophthalmidae) from Peru, with the description of three new species and a key to the Peruvian species. Herpetologica 60 (4):501-518 - get paper here
  • Kwet, A. 2007. Neue Schieneneidechse aus den Anden im Süden Perus. Draco 7 (27): 88 - get paper here
  • Stejneger, Leonhard 1913. Results of the Yale Peruvian Expedition of 1911.—Batrachians and reptiles. Proc. US. Natl. Mus. 45 (1992): 541-547 - get paper here
  • Werner,F. 1910. Über neue oder seltene Reptilien des Naturhistorischen Museums in Hamburg. ii. Eidechsen. Jahrb. Hamburg. Wiss. Anst., vol. 27 (1909), suppl. no. 2, 1910, pp. 1-46; reprinted: 1910, Mitteil. Naturhist. Mus. Hamburg, vol. 27: 205-) - get paper here
 
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