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Proctoporus pachyurus TSCHUDI, 1845

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Higher TaxaGymnophthalmidae (Cercosaurinae), Sauria, Gymnophthalmoidea, Squamata (lizards)
Subspecies 
Common NamesE: Tschudi's Lightbulb Lizard 
SynonymProctoporus pachyurus TSCHUDI 1845: 161
Proctoporus pachyurus — MÜLLER 1880
Proctoporus pachyurus – PETERS & DONOSO-BARROS 1970: 239
Proctoporus pachyurus — DOAN & SCHARGEL 2003 
DistributionPeru (Junin), elevation 2770–3800 m

Type locality: Valley of Río Chanchamayo, eastern slope of Andes, Peru  
Reproductionoviparous 
Types 
DiagnosisDiagnosis (genus Proctoporus sensu strictu): Proctoporus differs from members of subfamily Alopoglossinae by having its tongue covered in imbricate, scale-like papillae instead of oblique plicae (Harris, 1994), from Gymnophthalminae by having moveable eyelids, from Rhachisaurinae by having external ear openings and from Ecpelopinae by lacking prefrontal scales. Within Cercosaurinae (sensu Castoe et al., 2004), Proctoporus differs from all genera except Pholidobolus, Riama, and Petracola by lacking prefrontal scales. It differs from Riama by having an entire palpebral eye disc, from Petracola by having keeled dorsal scales and from Pholidobolus by having juxtaposed dorsal scales (after DOAN & CASTOE 2005. 
CommentType Species: Proctoporus pachyurus is the type species of the genus Proctoporus TSCHUDI 1845 sensu strictu (fide DOAN & CASTOE 2005).

Key: MAMANI et al. 2015 provide a key to all species of Proctoporus.

The P. pachyurus group (UZZELL 1958), unified by the presence of a single palpebral scale over the eye, a median occipital, and squarish pregular scales that do not form chevrons (Uzzell, 1970; Doan & Castoe, 2003), included P. bolivianus Werner, P. guentheri, and P. pachyurus. Doan & Castoe (2003) added two additional new species, P. sucullucu and P. unsaacae (after DOAN & CASTOE 2005). 
EtymologyNamed after Greek pachys (παχύς), thick; fat, plump + Greek oura (ουρα), tail. [?]. (from Esteban Lavilla, pers. comm., May 2024)

The genus was named after Greek proctos (πρωκτός), anus + Greek poros (πόρος), pore; crossing-place. ["...Pori anales per seriem semilunarem in ani margine posteriore dispositi…"]. 
References
  • CATENAZZI, A., LEHR, E. & VON MAY, R. 2013. The amphibians and reptiles of Manu National Park and its buffer zone, Amazon basin and eastern slopes of the Andes, Peru. Biota Neotropica 13 (4): 269-283
  • Doan, T. M. & Castoe, T.A. 2003. Using morphological and molecular evidence to infer species boundaries within Proctoporus bolivianus Werner (Squamata: Gymnopthalmidae). Herpetologica 59 (3): 432–449 - get paper here
  • Doan, T.F.; Castoe, T.A. & Arizábal Arriaga, W. 2005. Phylogenetic relationships of the genus Proctoporus sensu stricto (Squamata: Gymnophthalmidae), with a new species from Puno, Southeastern Peru. Herpetologica 61 (3): 325-336 - get paper here
  • Doan, T.M. 2003. A south-to-north biogeographic hypothesis for Andean speciation: evidence from the lizard genus Proctoporus (Reptilia, Gymnophthalmidae). Journal of Biogeography 30: 361–374 - get paper here
  • Goicoechea, Noemí; José M. Padial, Juan C. Chaparro, Santiago Castroviejo-Fisher, Ignacio De la Riva 2012. Molecular phylogenetics, species diversity, and biogeography of the Andean lizards of the genus Proctoporus (Squamata: Gymnophthalmidae). Molecular Phylogenetics and Evolution, 65 (3): 953–964 [print version published in 2013], http://dx.doi.org/10.1016/j.ympev.2012.08.017
  • Köhler, G. and E. Lehr 2004. Comments on Euspondylus and Proctoporus (Squamata: Gymnophthalmidae) from Peru, with the description of three new species and a key to the Peruvian species. Herpetologica 60 (4):501-518 - get paper here
  • Mamani L, Goicoechea N, Chaparro JC. 2015. A new species of Andean lizard Proctoporus (Squamata: Gymnophthalmidae) from montane forest of the Historic Sanctuary of Machu Picchu, Peru. Amphibian & Reptile Conservation 9 (1): 1–11 - get paper here
  • Müller, F. 1880. Erster Nachtrag zum Katalog der herpetologischen Sammlung des Basler Museums. Verh. Naturf. Ges. Basel 7: 120-165 - get paper here
  • Tschudi,J. J. von 1845. Reptilium conspectum quae in republica Peruana reperiuntur er pleraque observata vel collecta sunt in itenere. Archiv für Naturgeschichte 11 (1): 150-170 [reprint 1968, SSAR; sometimes cited as being published in 1846]. - get paper here
  • Uzzell, T.M. 1958. Teiid lizards related to Proctoporus luctuosus, with the description of a new species from Venezuela. Occasional Papers of the Museum of Zoology University of Michigan 597: 1-15 - get paper here
 
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