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Pseudocalotes drogon GRISMER, QUAH, WOOD, ANUAR, MUIN, DAVIS, MURDOCH, GRISMER, COTA & COBOS, 2016

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Drogon’s False Garden Lizard 
SynonymPseudocalotes drogon GRISMER, QUAH, WOOD, ANUAR, MUIN, DAVIS, MURDOCH, GRISMER, COTA & COBOS 2016
Pseudocalotes drogon — HONG et al. 2024 
DistributionPeninsular Malaysia (Pahang)

Type locality: 1 km south of Air Terjun Jeriau, Fraser’s Hill, Pahang, Peninsular Malaysia (3° 43.283 N 101° 43.035 E; 1066 m in elevation  
Reproductionoviparous. Each oviduct of the holotype contains a single large, oval egg with a thick fibrous shell that has not yet calcified (dimensions of egg from left oviduct 17.3 X 6.8 mm; clutch size two). The left ovary contains one large and subcircular yellow egg (largest diameter 4.8 mm) and four small cream-colored eggs (0.7–1.5 mm). 
TypesHolotype: LSUHC 12223, Adult male, collected on 24 March 2015 by L. Lee Grismer, Evan S. H. Quah, Perry L. Wood, Jr., Hayden R. Davis, Matthew L. Murdoch, Brando R. Burch, and Anthony J. Cobos at 2030 hrs. 
DiagnosisDiagnosis: Pseudocalotes drogon sp. nov. is differentiated from all other Psuedocalotes by having the combination of a flat rostrum; seven postrostrals; an interparietal; 11 circumorbitals; five canthals; 7–10 superciliaries; one scale between the rostral and nasal; nine supralabials; eight infralabials; 10 postnasal-suborbital scales; four postmentals; five or six sublabials; five or six chinshields; 47 smooth, wide, gular scales; weak transverse gular and antehumeral folds; two enlarged scales between the ear and eye; enlarged upper and lower posttemporals; a single enlarged supratympanic; no enlarged postrictals; three large scales bordering the dorsal margin of the ear opening; large pretympanic scales; eight scales in the nuchal crest not separated by a gap; enlarged vertebral scales extending to the tip of the tail; keeled and non-plate-like scales on flanks; 51 midbody scales; midventrals smaller than dorsals; 19 subdigital scales on the fourth finger; 23 subdigital scales on the fourth toe; preaxial scales on third toe enlarged and spinose; subdigital scales not unicarinate; HW/HL 0.52; HL/SVL 0.31; no elbow or knee patches; and a male dewlap color of lime-green bearing a central yellow spot. These characters or a subset of them are scored across all species in Table 3 (Grismer et al. 2016).


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CommentHabitat: Pseudocalotes drogon sp. nov. was collected at night at 2000 hrs while sleeping on a thin, horizontal branch of small tree in the vicinity of a small stream in hill dipterocarp forest (Fig. 7 in Grismer et al. 2016).

Coloration: The adult male was in the light color phase at the time of collection but became much darker during the day (Fig. 7 in Grismer et al. 2016).

Abundance: only known from the type specimen (Meiri et al. 2017). 
EtymologyThe specific epithet drogon refers to this species’ resemblance in form and color to the dark dragon, Drogon—one of three dragons born in the Dothraki Sea and commanded by Daenerys Targaryen—the Mother of Dragons—in George R. R. Martin’s fictional work Game of Thrones. 
References
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • GRISMER, L. LEE; EVAN S. H. QUAH, PERRY L. WOOD, JR., SHAHRUL ANUAR, ABDUL MUIN, HAYDEN R. DAVIS, MATTHEW L. MURDOCH, JESSE L. GRISMER, MICHAEL COTA & ANTHONY J. COBOS 2016. Dragons in the mist: three new species of Pseudocalotes Fitzinger (Squamata: Agamidae) from the sky island archipelago of Peninsular Malaysia. Zootaxa 4136 (3): 461–490 - get paper here
  • HONG, ZIJIA; M.S. SHAHRUL ANUAR, L. LEE GRISMER, & EVAN S. H. QUAH 2024. Expanded morphological description of the Drogon’s False Garden Lizard, Pseudocalotes drogon (Squamata: Agamidae) from Peninsular Malaysia. Zootaxa 5410 (4): 519–532 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Xu, Y., Gong, Y., Hou, M., Weng, S., Liu, S., Deng, J., ... & Peng, L. 2024. A New Species of the Genus Pseudocalotes (Squamata: Agamidae) from Southwest Yunnan, China. Animals, 14(6): 826 - get paper here
 
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