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Pseudocalotes guttalineatus HARVEY, HAMIDY, KURNIAWAN, SHANEY & SMITH, 2014

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Dash-lined False Garden Lizard 
SynonymPseudocalotes guttalineatus HARVEY, HAMIDY, KURNIAWAN, SHANEY & SMITH 2014 
DistributionIndonesia (S Sumatra)

Type locality: montane forest above Ngarip, Lampung, Sumatra, Indonesia, 5.28°S, 104.56° E, 1391 m elevation.  
Reproductionoviparous. The right oviduct of UTA 60501 contained two large, oval-shaped eggs (dimensions 14.6 X 6.9 mm and 14.2 X 7.1 mm; clutch size of four) already with tough fibrous shells, but no grossly evident calcification of the shell. In addition, this specimen’s right ovary contained two relatively large, yellow (diameters 4.7 mm) and 5 small, cream-colored (diameters 1–1.5 mm) eggs. The right oviduct of UTA 60540 was empty and highly convoluted, but not distended, suggesting that the specimen had not recently ovoposited. Its ovary contained one large, yellow, ovoid (4.8 X 3.7 mm) egg, a smaller circular egg (diameter 2.7), and 7 small cream-colored eggs (0.7–1.5 mm). 
TypesHolotype: MZB 9796, an adult female, collected 10 June 2013 by field parties from BC, MZB, UB, and UTA. Paratypes. Thirteen specimens (MZB 9652, 9765, 9792–9795, 9801, UTA 60536, 60540–60543, 60554) from the same mountain as the holotype, 1341–1521 m, collected 10–12 June 2013 by field parties from BC, MZB, UB, and UTA. Two specimens (UTA 60500–60501) from Maura Dua, Remanan Jaya, Gunung Pesagi (localy known as Masagi), Sumatera Selatan, Indonesia, 4.91°S, 104.13°E, 1574–1643 m, collected 17 June 2013 by field parties from BC, MZB, UB, and UTA. 
DiagnosisDiagnosis: A species of Pseudocalotes reaching at least 16 mm (95 mm SVL) and distinguished from congeners by the following combination of characters: (1) discrete interparietal absent (2) canthals 5–7, usually six; () enlarged, heavily keeled to subpyramidal nuchal, posttemporal, posttympanic, and postrictal scales present; (4) gulars medium-sized with sharp transition to smaller scales on well-developed gular pouch; (5) antehumeral skin fold absent; (6) 6–7 enlarged, heavily keeled scales in dorsolateral series between limbs, extending onto neck as short row of closely spaced scales; (7) scales on lower flanks feebly keeled and homogenous (8) 45–55 scales around midbody (9) dorsal crest of projecting scales, extending to tail of male and midbody of females; (10) ventrals smaller than dorsals; (11) subdigital lamellae at base of Toe III modified: prominent preaxial keels pointed; postaxial keels absent or greatly reduced; (12) dorsum green with dorsolateral series of pale grayish blue, heavily keeled scales; (1) venter light green, without pale ventrolateral stripe between limbs; (14) gular region with grayish blue lines evident when gular pouch extended; (15) tongue and floor of mouth yellow-orange, throat black.


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CommentSympatry: P. cybelidermus.

Although normally green in life, Pseudocalotes guttalineatus apparently can turn to various shades of brown, presumably in order to match its environment or temperament. 
EtymologyThe new name guttalineatus is an adjective derived from the Latin nouns gutta, meaning drop or spot, and lineatus, meaning lined. The new name refers to the dashed, bluish gray line on the neck and upper flanks of Pseudocalotes guttalineatus. 
References
  • HARVEY, MICHAEL B.; AMIR HAMIDY, NIA KURNIAWAN, KYLE SHANEY & ERIC N. SMITH 2014. Three new species of Pseudocalotes (Squamata: Agamidae) from southern Sumatra, Indonesia. Zootaxa 3841 (2): 211–238 - get paper here
  • HARVEY, MICHAEL B.; KYLE SHANEY, AMIR HAMIDY, NIA KURNIAWAN, ERIC N. SMITH 2017. A new species of Pseudocalotes (Squamata: Agamidae) from the Bukit Barisan Range of Sumatra with an Estimation of its phylogeny. Zootaxa 4276 (2): 215–232 - get paper here
  • Xu, Y., Gong, Y., Hou, M., Weng, S., Liu, S., Deng, J., ... & Peng, L. 2024. A New Species of the Genus Pseudocalotes (Squamata: Agamidae) from Southwest Yunnan, China. Animals, 14(6): 826 - get paper here
 
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