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Pseudocalotes rhaegal GRISMER, QUAH, WOOD, ANUAR, MUIN, DAVIS, MURDOCH, GRISMER, COTA & COBOS, 2016

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Rhaegal’s False Garden Lizard 
SynonymPseudocalotes rhaegal GRISMER, QUAH, WOOD, ANUAR, MUIN, DAVIS, MURDOCH, GRISMER, COTA & COBOS 2016 
DistributionPeninsular Malaysia (Pahang)

Type locality: Robinson Falls, Cameron Highlands, Pahang, Peninsular Malaysia (04° 43.283 N 101° 23.129 E; 1411 m in elevation  
Reproductionoviparous. All were gravid females collected during mid-March or mid- September, indicating that this species may breed year-round (Grismer et al. 2016). 
TypesHolotype: LSUHC 12178, Adult female, collected on 18 March 2015 by L. Lee Grismer, Evan S. H. Quah, Shahrul Anuar, Mohd A. Muin, Perry L. Wood, Jr., Hayden R. Davis, Matthew L. Murdoch, Brandon R. Burch, and Anthony J. Cobos at 2030 hrs. Paratypes. Adult female LSUHC 12179 bears the same locality collecting data as the holotype. Adult female LSUHC 12000 bears the same collecting locality and collectors but was collected on 4 September 2014. 
DiagnosisDiagnosis: Pseudocalotes rhaegal sp. nov. is differentiated from all other Psuedocalotes by having the combination of a convex rostrum; 6–8 postrostrals; an interparietal; nine or 10 circumorbitals; five canthals; 7–10 superciliaries; one or two scales between the rostral and nasal scales; eight or nine supralabials; seven or eight infralabials; 11 or 12 postnasal-suborbital scales; four postmentals; 4–6 chinshields; 40–45 smooth, wide, gular scales; no transverse gular fold; weak antehumeral fold; three or four enlarged scales between the ear and eye; an enlarged upper and lower posttemporal; an enlarged supratympanic; no enlarged postrictals; no large scales bordering the upper margin of the ear opening or in the pretympanic region; 6–8 enlarged nuchal crest scales not separated by a gap; enlarged vertebral scales extending to base of tail; weakly keeled, non-plate-like scales on flanks; 52–58 scales around midbody; midventrals smaller than dorsals; 19–21 subdigital lamellae on fourth finger; 22–26 subdigital lamellae on fourth toe; preaxial scales on third toe enlarged and rounded; subdigital lamellae not unicarinate; HW/HL 0.50–0.54; HL/SVL 0.28–0.30; no elbow or knee patches; and female dewlap bearing a purple base. These characters or a subset of them are scored across all species in Table 3 (Grismer et al. 2016).


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CommentHabitat: All Pseudocalotes rhaegal sp. nov. were collected at night between at 2000 and 2300 hrs while sleeping on thin, horizontal branches of small trees in the vicinity of a river flowing through the hill dipterocarp forest at Robinson Falls (Grismer et al. 2016).

Abundance: only known from its original description (Meiri et al. 2017). 
EtymologyThe specific epithet rhaegal refers to this species’ resemblance in form and color to the greenish dragon, Rhaegal—one of three dragons born in the Dothraki Sea and commanded by Daenerys Targaryen—the Mother of Dragons—in George R. R. Martin’s fictional work Game of Thrones. 
References
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • GRISMER, L. LEE; EVAN S. H. QUAH, PERRY L. WOOD, JR., SHAHRUL ANUAR, ABDUL MUIN, HAYDEN R. DAVIS, MATTHEW L. MURDOCH, JESSE L. GRISMER, MICHAEL COTA & ANTHONY J. COBOS 2016. Dragons in the mist: three new species of Pseudocalotes Fitzinger (Squamata: Agamidae) from the sky island archipelago of Peninsular Malaysia. Zootaxa 4136 (3): 461–490 - get paper here
  • HONG, ZIJIA; M.S. SHAHRUL ANUAR, L. LEE GRISMER, & EVAN S. H. QUAH 2024. Expanded morphological description of the Drogon’s False Garden Lizard, Pseudocalotes drogon (Squamata: Agamidae) from Peninsular Malaysia. Zootaxa 5410 (4): 519–532 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Xu, Y., Gong, Y., Hou, M., Weng, S., Liu, S., Deng, J., ... & Peng, L. 2024. A New Species of the Genus Pseudocalotes (Squamata: Agamidae) from Southwest Yunnan, China. Animals, 14(6): 826 - get paper here
 
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