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Pseudocalotes viserion GRISMER, QUAH, WOOD, ANUAR, MUIN, DAVIS, MURDOCH, GRISMER, COTA & COBOS, 2016

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Viserion’s False Garden Lizard 
SynonymPseudocalotes viserion GRISMER, QUAH, WOOD, ANUAR, MUIN, DAVIS, MURDOCH, GRISMER, COTA & COBOS 2016 
DistributionPeninsular Malaysia (Pahang)

Type locality: crossing the radar tower road at Ulu Kali at Genting Highlands, Pahang, Peninsular Malaysia (03° 26.166 N 101° 47.021 E; 1754 m in elevation  
Reproductionoviparous. The holotype was a gravid female that had just ovulated indicating that the reproductive season of this species extends through March. 
TypesHolotype: LSUHC 12227, Adult female, collected on 26 March 2015 by Evan S. H. Quah at 1000 hrs. Paratype. Adult male LSUHC 12141 found dead on the radar tower road at Ulu Kali at Genting Highlands, Pahang, Peninsular Malaysia (03° 26.12 N 101° 47.345 E; 1750 m in elevation) by L. Lee Grismer, Perry L. Wood, Jr., Hayden R. Davis, Matthew L. Murdoch, Brandon R. Burch, and Anthony J. Cobos. 
DiagnosisDiagnosis: Pseudocalotes viserion sp. nov. can be separated from all other species of Psuedocalotes by having a combination of three postrostrals; 10 circumorbitals; four or five canthals; 5–7 superciliaries; rostral and nasal in contact; supralabials contacting nasal; six or seven supralabials; seven or eight infralabials; two or three postmentals; three enlarged chinshields; 47 or 48 smooth, flat, gular scales; weak transverse gular and antehumeral folds; two enlarged scales between the ear and eye; an enlarged upper and lower posttemporal; an enlarged supratympanic; no enlarged postrictals; 7–9 nuchal crest scales lacking gaps and not extending beyond midbody; weakly keeled and plate-like scales on flanks; 35–38 scales around midbody; ventrals smaller than dorsals; 22 or 23 subdigital lamellae on fourth finger; 26 or 27 subdigital lamellae on fourth toe; preaxial scales on third not modified; subdigital lamellae not unicarinate; HW/HL 0.62; no white marking below the eye; dewlap in males yellow; and no elbow or knee patches. These characters or a subset of them are scored across all species in Table 3 (Grismer et al. 2016).


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CommentHabitat: The holotype (LSUHC 12227) was collected during mid-morning at 1000 hrs under broken cloud cover as it was crossing the radar tower road at Ulu Kali at Genting Highlands. The paratype (LSUHC 12141) was found freshly killed on the same road at 1500 hrs. This short stretch of road runs along the crest of Ulu Kali and winds through a mossy cloud forest between 1,700 and 1,800 m in elevation (Fig. 11 in Grismer et al. 2016).

Abundance: only known from its original description (Meiri et al. 2017). 
EtymologyThe specific epithet viserion refers to this species’ resemblance in form and color to the yellowish dragon, Viserion—one of three dragons born in the Dothraki Sea and commanded by Daenerys Targaryen—the Mother of Dragons—in George R. R. Martin’s fictional work Game of Thrones. 
References
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • GRISMER, L. LEE; EVAN S. H. QUAH, PERRY L. WOOD, JR., SHAHRUL ANUAR, ABDUL MUIN, HAYDEN R. DAVIS, MATTHEW L. MURDOCH, JESSE L. GRISMER, MICHAEL COTA & ANTHONY J. COBOS 2016. Dragons in the mist: three new species of Pseudocalotes Fitzinger (Squamata: Agamidae) from the sky island archipelago of Peninsular Malaysia. Zootaxa 4136 (3): 461–490 - get paper here
  • HONG, ZIJIA; M.S. SHAHRUL ANUAR, L. LEE GRISMER, & EVAN S. H. QUAH 2024. Expanded morphological description of the Drogon’s False Garden Lizard, Pseudocalotes drogon (Squamata: Agamidae) from Peninsular Malaysia. Zootaxa 5410 (4): 519–532 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Miranda JP, Costa JCL, Rocha CFD 2012. Reptiles from Lençóis Maranhenses National Park, Maranhão, northeastern Brazil. ZooKeys 246: 51–68 - get paper here
  • Xu, Y., Gong, Y., Hou, M., Weng, S., Liu, S., Deng, J., ... & Peng, L. 2024. A New Species of the Genus Pseudocalotes (Squamata: Agamidae) from Southwest Yunnan, China. Animals, 14(6): 826 - get paper here
 
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