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Pseudocalotes kingdonwardi (SMITH, 1935)

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Common NamesE: Kingdonward's Bloodsucker
kaulbacki: Smith's Japalure
Chinese: 西藏拟树蜥 
SynonymCalotes kingdon-wardi SMITH 1935: 204
Japalura kaulbacki SMITH 1937: 755
Japalura kaulbacki — WERMUTH 1967: 66
Calotes kingdonwardi — WERMUTH 1967: 39
Calotes kingdonwardi — BARTS & WILMS 2003
Japalura kaulbacki — BARTS & WILMS 2003
Calotes kingdonwardi kingdonwardi — MANTHEY 2008: 72
Pseudocalotes kingdonwardi — MAHONY 2010
Pseudocalotes kaulbacki — VENUGOPAL 2010 
DistributionChina (W Yunnan)
Border area between Tibet and Burma (= Myanmar)

Type locality: Adung Valley (7000 f.), on the Burma-Tibetan border

kaulbacki: N Myanmar (= Burma); Type locality: Nam Tamai Valley, Myanmar (= Burma)-Tibet border (elevation: 3500 feet)  
TypesHolotype: BMNH 1946.8.11.17 (was BMNH 1932.6.8.8 fide SMITH 1935, MAHONY 2010)
Holotype: BMNH 1936.7.4.19/1946.9.4.72 [kaulbacki] 
Diagnosis“Vindum et al. (2003) described one minor differences between the type specimens of Calotes kingdonwardi bapoensis and the original description of C. kingdonwardi, namely; “scales in C. kingdonwardi pointing backwards and downwards except for upper two to three rows, which point slightly upwards (Smith 1935) or straight backwards [KIZ specimens=type series]”. The upper scale rows vary between straight back and slightly upwards on examined adult P. kaulbacki new comb. (=P. kingdonwardi new comb.), indicating likely intrapopulational variation. Furthermore the plates provided of the type specimens of Calotes kingdonwardi bapoensis (Yang et al. 1979) clearly show the enlarged subimbricate lateral gular scales decreasing in size considerably towards the centre of the throat. This character is identical to that of most species of Pseudocalotes (i.e., P. austeniana new comb., P. brevipes, P. dringi, P. flavigula, P. floweri, P. kakhienensis new comb., P. khaonanensis, P. kingdonwardi new comb., P. microlepis, P. poilani, P. saravacensis), but not present on any of the examined Japalura species.” [from MAHONY 2010]. 
CommentSynonymy: MAHONY 2010 synonymized Japalura kaulbacki with Calotes kingdonwardi. He also treted bapoensis as a synonym of kingdonwardi, given that “the English extract of the original description does not provide any characters that differ from P. kingdonwardi”.

The taxonomic status of C. k. bapoensis should be assessed relative to P. kingdonwardi new comb. as redefined here. If it is found to represent a valid species, then it should be referred to as Pseudocalotes bapoensis (Yang & Su, 1979) new comb.” [from MAHONY 2010]. 
EtymologyNamed after the collector, Captain Kingdon-Ward (1885–1956). 
  • Barts, M. & Wilms, T. 2003. Die Agamen der Welt. Draco 4 (14): 4-23 - get paper here
  • Hallermann, J. 2000. A new species of Calotes from the Moluccas (Indonesia) with notes on the biogeogreaphy of the genus (Sauria: Agamidae). Bonner Zoologische Beiträge 49 (1-2): 155-163 - get paper here
  • HARVEY, MICHAEL B.; AMIR HAMIDY, NIA KURNIAWAN, KYLE SHANEY & ERIC N. SMITH 2014. Three new species of Pseudocalotes (Squamata: Agamidae) from southern Sumatra, Indonesia. Zootaxa 3841 (2): 211–238 - get paper here
  • MAHONY, STEPHEN 2010. Systematic and taxomonic revaluation of four little known Asian agamid species, Calotes kingdonwardi Smith, 1935, Japalura kaulbacki Smith, 1937, Salea kakhienensis Anderson, 1879 and the monotypic genus Mictopholis Smith, 1935 (Reptilia: Agamidae). Zootaxa 2514: 1–23 - get paper here
  • Manthey U 2008. Agamid lizards of Southern Asia, Draconinae 1. Terralog 7, 160 pp.
  • Smith, M.A. 1935. The fauna of British India, including Ceylon and Burma. Reptiles and Amphibia, Vol. II. Sauria. Taylor and Francis, London, 440 pp.
  • Smith, M.A. 1937. Description of a new species of agamid lizard from Upper Burma. J. Bombay Nat. Hist. Soc. 39: 755 - get paper here
  • Smith, M.A. 1940. The Amphibians and Reptiles obtained by Mr. Ronald Kaulback in Upper Burma. Records of the Indian Museum 42: 465-486
  • Venugopal, P.D. 2010. An updated and annotated list of Indian lizards (Reptilia: Sauria) based on a review of distribution records and checklists of Indian reptiles. Journal of Threatened Taxa 2 (3): 725-738. - get paper here
  • Vindum, J. V., Htun Win, Thin Thin, Kyi Soe Lwin, Awan Khwi Shein & Hla Tun 2003. A new Calotes (Squamata: Agamidae) from the Indo-Burman range of western Myanmar (Burma). Proc. Cal. Acad. Sci. 54(1): 1-16 - get paper here
  • Wang, Kai; Jinlong Ren, Hongman Chen, Zhitong Lyu, Xianguang Guo Ke Jiang, Jinmin Chen, Jiatang Li, Peng Guo, Yingyong Wang, Jing Che 2020. The updated checklists of amphibians and reptiles of China. Biodiversity Science 28 (2): 189-218 - get paper here
  • Yang D T; Su C Y; Li S M 1979. New species and new subspecies of amphibians and reptiles from Gaoligong Shan, Yunnan. Acta Zootaxonomica Sinica 4(2) 1979: 185-188
  • Zug, G. R., H. H. K. Brown, J. A. Schulte II, and J. V. Vindum 2006. Systematics of the Garden Lizards, Calotes versicolor Group (Reptilia, Squamata, Agamidae), in Myanmar: Central Dry Zone Populations. Proc. Cal. Acad. Sci. 57 (2):35-68. - get paper here
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