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Pseudogonatodes furvus RUTHVEN, 1915

IUCN Red List - Pseudogonatodes furvus - Data Deficient, DD

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Higher TaxaSphaerodactylidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Colombian Clawed Gecko 
SynonymPseudogonatodes furvus RUTHVEN 1915
Pseudogonatodes furvus — PETERS & DONOSO-BARROS 1970: 242
Pseudogonatodes furvus — KLUGE 1993
Pseudogonatodes furvus — RÖSLER 2000: 106 
DistributionColombia (Magdalena)

Type locality: San Lorenzo, Santa Marta Mountains, 5000 ft., restricted to “Vereda Bella Vista, Sierra Nevada de Santa Marta”, corregimiento de Minca, Distrito de Santa Marta, Magdalena, Colombia by Montes-Correa et al. 2021.  
Reproductionoviparous 
TypesHolotype: UMMZ 47782, adult male collected July 20, 1913 by Frederick M. Gaige. Other specimens: MCZ, ICN-R, CBUMAG 
DiagnosisDiagnosis (genus, tentatively); a group that was characterized by the presence of granular dorsal scales, long digits with many subdigital lamellae (8–15 lamellae under fourth toe), and the absence of third enlarged lamella in fourth toe (Huey & Dixon 1970, Montes-Correa et al. 2021).

Definition. Pseudogonatodes furvus can be distinguished from all of its congeners by the following combination of characters (Figure 1): (1) maximum SVL of 45 mm for males and 43.8 mm for females; (2) dorsal scales homogenous, granular, subimbricate, and larger than occipital scales; (3) 83–104 scales around midbody; (4) posterior edge of rostral indented by three postrostrals, with a median cleft; (5) 4–5 supralabials; (6) 4–5 infralabials; (7) 7–10 loreals; (8) posterior edge of mental scale with two median clefts (W-shaped), indented by (9) 2–4 postmentals, which are larger than subsequent scales; (10) 44–67 ventrals in straight line between anterior level of forelimbs and border of cloaca, (11) 34–39 ventrals between anterior levels of forelimbs and hindlimbs; (12) long digits lacking expanded subdigital third lamella, (13) 11–13 subdigital lamellae under finger IV and (14) 11–15 subdigital lamellae under toe IV; (15) subcaudal pattern with sequence of moderately to heavily enlarged medial scales in contact laterally with one to two scales (Montes-Correa et al. 2021: 44).

Comparisons. A summary of morphological characters of the genus Pseudogonatodes is given in Table 3. Pseudogonatodes furvus is quickly distinguished from all members of the Group I (P. guianensis and P. lunulatus) sensu Huey & Dixon (1970) by the following characters: absence of enlarged third lamellae under fourth toe (present in Group I); posterior edge of mental with two median clefts (W-shaped) and postmentals larger than following scales (posterior border of mental slightly concave to heavily concave [U-shaped], postmentals equal in size to following scales); and sole of foot with homogeneous squamation (heterogeneous squamation in P. lunulatus). Regarding the only member of the Group II (P. barbouri), P. furvus is distinguishable by the absence of enlarged third subdigital lamellae under the fourth toe (present in P. barbouri); dorsals small, granular, conical, and sub-imbricated (large, flat, and imbricate). From Group III, P. furvus differs by having posterior edge of mental with two median clefts (U-shaped without clefts in P. peruvianus Huey & Dixon 1970, V-shaped with a median cleft in P. gasconi Avila-Pires & Hoogmoed 2000); two to four postmentals larger than following scales (six equal in size to following scales for P. gasconi, six to eight slightly larger than following scales for P. manessi Avila-Pires & Hoogmoed 2000); subcaudal pattern with sequence of moderately to heavily enlarged midventral scales in contact laterally with one to two scales (no enlarged midventral subcaudal scales in P. gasconi); and seven to ten loreals (six in P. peruvianus) (Montes-Correa et al. 2021: 44). 
CommentConservation: the species is only known from a few specimens from 3 localities.

Distribution: erroneously reported from the Villavicencio region of Colombia (PETERS & OREJAS-MIRANDA 1970). For maps of species distributions see Vanzolini 1968: 92 (Map 3) and Montes-Correa et al. 2021: 53 (Fig. 5).

Type species: Pseudogonatodes furvus RUTHVEN 1915 is the type species of the genus Pseudogonatodes RUTHVEN 1915.

Key: Huey & Dixon 1970 present a key to the species of Pseudogonatodes. 
References
  • Avila-Pires, T.C.S. & Hoogmoed,M.S. 2000. On two new species of Pseudogonatodes Ruthven, 1915 (Reptilia: Squamata: Gekkonidae), with remarks on the distribution of some other sphaerodactyl lizards. Zoologische Mededelingen 73 (12): 209-223 - get paper here
  • Carvajal-Cogollo, J.E.; L.E. Rojas-Murcia. & G. Cárdenas-Arévalo 2020. Reptiles del Caribe colombiano/ Reptiles of the Colombian Caribbean. Tunja: Editorial UPTC, 268 pp. - get paper here
  • Huey, R. B., and J. R. Dixon. 1970. A new Pseudogonatodes from Peru with remarks on other species of the genus. Copeia 1970 (3): 538—542 - get paper here
  • MONTES-CORREA, ANDRÉS CAMILO; LILIANA P. SABOYÁ-ACOSTA, JUAN DAVID JIMÉNEZ-BOLAÑO, TEDDY ANGARITA-SIERRA, VLADIMIR BRICEÑO-PÉREZ, SAMUEL NÚÑEZ, JUAN MANUEL RENJIFO, WALTER E. SCHARGEL, JUAN D. DAZA, MARINUS S. HOOGMOED 2021. Extended diagnosis of the type species of Pseudogonatodes Ruthven 1915 (Gekkota: Sphaerodactylidae). Zootaxa 4915 (1): 041–059 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Ruthven, Alexander G. 1915. Description of a new genus and species of lizard of the family Gekkonidæ. Occasional Papers of the Museum of Zoology, University of Michigan (19): 1-3 - get paper here
 
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