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Pseudotrapelus neumanni (TORNIER, 1905)

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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Neumann’s Agama 
SynonymAgama neumanni TORNIER 1905: 384
Pseudotrapelus neumanni — MELNIKOV et al. 2013
Pseudotrapelus neumanni — MELNIKOV et al. 2015
Pseudotrapelus neumanni — TAMAR et al. 2016 
DistributionS Yemen, SW Saudi Arabia (Asir Mountains)

Type locality: Haithalhin, Lahadj (paratype label in NMW)  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: ZMB 42952; paratypes: ZMB 27418, 37225, 54522, and 54523, NMW 23341 
DiagnosisDiagnosis: “Pseudotrapelus neumanni can be distinguished, as this is the only Pseudotrapelus species with enlarged scales on the dorsal parts of head and body.” (MELNIKOV et al. 2015: 59). 
CommentSynonymy: ARNOLD 1980 synonymized A. neumanni with A. sinaitus. MELNIKOV et al. 2013, 2015 and TAMAR et al. 2016 consider Pseudotrapelus neumanni as a valid species. However, Melnikov’s description/diagnosis seems rather vague, without any illustrations or phylogenetic analysis.

Distribution: see maps in TAMAR et al. 2016: Fig. 1, Tamar et al. 2019: Fig. 1, Tamar et al. 2023: 1034 (Fig. 1).

Reference images: see Uetz et al. 2024 for high-resolution reference images for this species. 
EtymologyNamed after Oscar Neumann (3 September 1867–17 May 1946) who undertook in the late 19th and early 20th centuries influential expeditions to East Africa, Somalia, Ethiopia, and Sudan. Neumann worked voluntarily for over 40 years in the Museum für Naturkunde, Berlin, but was forced by the Nazi government to leave Germany in 1941. 
References
  • Arnold E N 1980. The scientific results of the Oman flora and fauna survey 1977 (Dhofar). The reptiles and amphibians of Dhofar, southern Arabia. JOURNAL OF OMAN STUDIES SPECIAL REPORT (No. 2): 273-332 - get paper here
  • Gemel, R.; G. Gassner & S. Schweiger 2019. Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien – 2018. Ann. Naturhist. Mus. Wien, B 121: 33–248
  • Melnikov, D.A. and E.N. Melnikova 2013. Taxonomic diversity of the genus Pseudotrapelus (Agamidae; Sauria) and biogeography of Afro-Arabia. [in Russian, with English Abstract]. Proceedings of the Zoological Institute of the Russian Academy of Sciences 317 (4): 438–451 - get paper here
  • Melnikov, Daniel; Jan Śmiełowski, Ekaterina Melnikova, Roman Nazarov, Natalia B. Ananjeva 2015. Red’n’blues: A New Species of Pseudotrapelus (Agamidae, Sauria) from Sudan, Africa. Russ. J. Herpetol. 22 (1): 53-60 - get paper here
  • Melnikova, Ekaterina; Daniel Melnikov, Natalia Ananjeva 2015. Mitochondrial and Nuclear Gene Phylogeny of Pseudotrapelus (Sauria: Agamidae) with Considerations on the Hybridization and mtDNA Introgression between Species. Russ. J. Herpetol. 22 (1): 61-67 - get paper here
  • Tamar, K., Chirio, L., Shobrak, M., Busais, S. & Carranza, S. 2019. Using multilocus approach to uncover cryptic diversity within Pseudotrapelus lizards from Saudi Arabia. Saudi J. Biol. Sci. (2019). doi: - get paper here
  • Tamar, K., Uvizl, M., Shobrak, M., Almutairi, M., Busais, S., AlGethami, R. H. M., ... & Šmíd, J. 2023. A new species of Pseudotrapelus (Reptilia: Squamata: Agamidae) from Central Arabia. Vertebrate Zoology 73, 1033-1045 - get paper here
  • Tamar, Karin; Sebastian Scholz, Pierre-André Crochet, Philippe Geniez, Shai Meiri, Andreas Schmitz, Thomas Wilms, Salvador Carranza 2016. Evolution around the Red Sea: Systematics and biogeography of the agamid genus Pseudotrapelus (Squamata: Agamidae) from North Africa and Arabia. Molecular Phylogenetics and Evolution 97 (2016) 55–68 - get paper here
  • Tornier, G. 1905. Schildkröten und Eidechsen aus Nord-Ost Afrika und Arabien. Aus Carlo v. Erlanger’s und Oscar Neumann’s Forschungsreise. Zool. Jahrb. Syst. 22: 365-388 - get paper here
  • Uetz, P.H.; Patel, M.; Gbadamosi, Z.; Nguyen, A.; Shoope, S. 2024. A Reference Database of Reptile Images. Taxonomy 4: 723–732 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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