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Ptyas nigromarginata (BLYTH, 1854)

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Green Rat Snake, Black-bordered Rat Snake
G: Grüne Gekielte Rattennatter
Chinese: 黑线乌梢蛇 
SynonymColuber nigromarginatus BLYTH 1854
Zaocys nigromarginatus — GÜNTHER 1864
Zaocys nigromarginatus — BOULENGER 1893: 376
Zaocys nigromarginatus — WALL 1908: 325
Zaocys dhumnades nigromarginatus — ANGEL & BOURRET 1933
Zaocys nigromarginatus — SMITH 1943: 165
Zaocys nigromarginatus — SWAN & LEVITON 1962: 116
Coluber nigromarginatus — DAS 1996: 58
Ptyas nigromarginata — DAVID & DAS 2004
Zaocys nigromarginatus — SHARMA 2004
Ptyas nigromarginata — WHITAKER & CAPTAIN 2004
Ptyas nigromarginatus — PYRON & BURBRINK 2013
Ptyas nigromarginata — WALLACH et al. 2014: 617 
DistributionNepal, India (Darjelling, Sikkim, Assam ?, Meghalaya, Arunachal Pradesh, Nagaland), N Myanmar (= Burma), N Vietnam (Lao Cai Province, Lang Son, Ha Tay, Kon Tum Province, Ha Giang), Thailand (Nan), Bhutan, Bangladesh,
China (Guizhou, Yunnan, SW Sichuan, SE Xizang = Tibet), elevation 500-2290 m.

Type locality: Vicinity of Darjeeling, India  
Reproductionoviparous. 
TypesHolotype: ZSI 7343 
Diagnosis 
CommentSynonymy: Lazell et al. (1991) and Lazell (1998) formally synonymized Ptyas and Coluber but at least the latter doesn’t explicitely mention Ptyas nigromarginatus.

Similar species: juveniles are strikingly similar to the juveniles of the species P. dhumnades.

Distribution: Reports from Afghanistan are most likely in error (Vogel & Hauser 2013). These authors listed many localities. 
References
  • Bai, Linzhuang and Kai Wang. 2015. Ptyas nigromarginata (green ratsnake) diet. Herpetological Review 46 (3): 453 - get paper here
  • Bauer A M; Gunther R 1992. A preliminary report on the reptile fauna of the Kingdom of Bhutan with the description of a new species of scincid lizard (Reptilia: Scincidae). Asiatic Herpetological Research 4: 23-36 - get paper here
  • Blyth, EDWARD. 1855. Notices and descriptions of various reptiles, new or little known [part 2]. J. Asiatic Soc. Bengal, Calcutta, 23 (3): 287-302 [1854] - get paper here
  • Boulenger, G.A. 1893. Catalogue of the snakes in the British Museum (Nat. Hist.) I. London (Taylor & Francis), 448 pp. - get paper here
  • Das, I. 2012. A Naturalist's Guide to the Snakes of South-East Asia: Malaysia, Singapore, Thailand, Myanmar, Borneo, Sumatra, Java and Bali. Oxford J, ohn Beaufoy Publishing - get paper here
  • David, P. & I. Das 2004. On the grammar of the gender of Ptyas Fitzinger, 1843 (Serpentes: Colubridae). Hamadryad 28 (1&2): 113-116
  • Dowling, H.G., & Jenner, J.V. 1988. Snakes of Burma: checklist of reported species and bibliography. Smithsonian Herp. Inf. Serv. (76): 19 pp. - get paper here
  • Kästle , W., Rai, K. & Schleich, H.H. 2013. FIELD GUIDE to Amphibians and Reptiles of Nepal. ARCO-Nepal e.V., 625 pp. - get paper here
  • Lazell,J.D. 1998. Morphology and the status of the snake genus "Ptyas". Herpetological Review 29 (3): 134 - get paper here
  • Lazell,J.D., Keirans,J.E. & Samuelson,G.A. 1991. The Sulawesi black racer, Coluber (Ptyas) dipsas, and a remarkable ectoparasitic aggregation. Pacific Science 45 (4): 355-361
  • Lenz, Norbert 2012. Von Schmetterlingen und Donnerdrachen - Natur und Kultur in Bhutan. Karlsruher Naturhefte 4, Naturkundemuseum Karlsruhe, 124 pp.
  • Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • O’Shea, M. 2018. The Book of Snakes. Ivy Press / Quarto Publishing, London, - get paper here
  • Pyron, R. Alexander; Frank T. Burbrink 2013. Early origin of viviparity and multiple reversions to oviparity in squamate reptiles. Ecology Letters 17 (1): 13–21 (published online 2013, in print 2014), DOI: 10.1111/ele.12168 - get paper here
  • Sharma, R. C. 2004. Handbook Indian Snakes. AKHIL BOOKS, New Delhi, 292 pp.
  • Smith, M.A. 1943. The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. 3 (Serpentes). Taylor and Francis, London. 583 pp.
  • Swan, L.W., & Leviton, A.E. 1962. The herpetology of Nepal: a history, check list, and zoogeographical analysis of the herpetofauna. Proc. Cal. Acad. Sci. 32 (6) (4.s.): 103-147. - get paper here
  • Tshewang, S. & L. Letro 2018. The herpetofauna of Jigme Singye Wangchuck National Park in central Bhutan: status, distribution and new records. Journal of Threatened Taxa 10(11): 12489–12498; - get paper here
  • VOGEL, Gernot and Sjon HAUSER 2013. Addition of Ptyas nigromarginata (Blyth, 1854) (Squamata: Colubridae) to the Snake Fauna of Thailand with Preliminary Remarks on Its Distribution. Asian Herpetological Research 4 (3): 166–181
  • Wall,F. 1908. Notes on a collection of snakes from the Khasi Hills, Assam. J. Bombay nat. Hist. Soc. 18: 312-337 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Wang, Kai; Jinlong Ren, Hongman Chen, Zhitong Lyu, Xianguang Guo Ke Jiang, Jinmin Chen, Jiatang Li, Peng Guo, Yingyong Wang, Jing Che 2020. The updated checklists of amphibians and reptiles of China. Biodiversity Science 28 (2): 189-218 - get paper here
  • Wangyal, Jigme Tshelthrim 2019. The status of herpetofauna of Bhutan. District Forest Office, District Administration, Trashigang, Bhutan, 20-39 - get paper here
  • Whitaker, Romulus and Ashok Captain 2004. Snakes of India. Draco Books, 500 pp., reprinted 2007 - get paper here
  • Yang, J.-H., X.-Y. Huang, J.-F. Ye, S.-P. Yang, X.-.C. Zhang & B.P.-L. Chan 2019. A report on the herpetofauna of Tengchong Section of Gaoligongshan National Nature Reserve, China. Journal of Threatened Taxa 11(11): 14434–14451 - get paper here
 
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