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Ptyctolaemus gularis (PETERS, 1864)

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Green Fan-throated lizard
G: Kehlfaltenagame
Chinese: 喉褶蜥 
SynonymOtocryptis (Ptyctolaemus) gularis PETERS 1864: 386
Ptyctolaemus gularis — BOULENGER 1885: 273
Ptyctolaemus gularis — ANNANDALE 1905
Ptyctolaemus gularis — HORA 1926
Ptyctolaemus gularis — SMITH 1935: 149
Ptyctolaemus gularis — SMITH 1940: 476
Ptyctolaemus gularis — DAS 1996: 45
Ptyctolaemus gularis — MAHONY et al. 2009 
DistributionNE India (Assam; Khasi hills; Arunachal Pradesh, Mizoram, Nagaland), Bangladesh,
China (Xizang = Tibet), Myanmar (Kachin, Chin)

Type locality: “Calcutta”; in error fide ZHAO & ADLER 1993.  
Reproductionoviparous 
TypesHolotype: ZMB 5004; additional material: ZMB 31134-31135, from Margherita, in Patkai Mountains, Upper Assam (fide ANANJEVA & STUART 2001). 
DiagnosisDiagnosis (genus): see P. collicristatus. 
CommentType species: Otocryptis (Ptyctolaemus) gularis PETERS 1864 is the type species of the genus Ptyctolaemus PETERS 1864.

Distribution: possibly in Bhutan (Lenz 2012). Possibly in Yunnan (China) but records from there are questionable (Wang et al. 2022).

Reference images: see Uetz et al. 2024 for high-resolution reference images for this species. 
References
  • Ahsan, M. Farid; Asmat, G.S.M. & Chakma, S. 2008. A Second Record of Ptyctolaemus gularis (Peters, 1864) from Bangladesh. Asiatic Herpetological Research 11: 10-12 - get paper here
  • ANANJEVA, N. B. & STUART. B.L. 2001. The agamid lizard Ptyctolaemus phuwuanensis MANTHEY AND NABHITABHATA, 1991 from Thailand and Laos represents a new genus. Russ. J. Herpetol. 8 (3): 165-170 - get paper here
  • Annandale, Nelson 1905. Contributions to Oriental Herpetology. Suppl. II. Notes on the Oriental lizards in the Indian Museum, with a list of the species recorded from British India and Ceylon. J. Proc. Asiat. Soc. Bengal (2) 1: 81-93 - get paper here
  • Bhupathy, S., Ramesh Kumar, S., Paramanandham, J., Thirumalainathan, P. and Pranjit Kumar Sarma 2013. Conservation of reptiles in Nagaland, India. In: K. K. Singh et al. (editors), Bioresources and Traditional Knowledge of Northeast India, Mizo Post-Graduate Science Society, pp. 181-186
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Boulenger, George A. 1890. The Fauna of British India, Including Ceylon and Burma. Reptilia and Batrachia. Taylor & Francis, London, xviii, 541 pp. - get paper here
  • Das, Abhijit 2016. Addition to the Herpetofauna of Royal Manas National Park, Bhutan, with six new country records. Herpetology Notes 9: 261-278 - get paper here
  • Das, Indraneil & Abhijit Das 2017. A Naturalist’s Guide to the Reptiles of India, Bangladesh, Bhutan, Nepal, Pakistan and Sri Lanka. John Beaufoy Publishing Ltd., Oxford, 176 pp.
  • DECEMSON, H.T. & H.T. LALREMSANGA. 2022. PTYCTOLAEMUS GULARIS (Green Fan throated Lizard). WATER INTAKE. Herpetological Review 53 (2): 330.
  • Hakim, J., Trageser, S. J., Ghose, A., Rashid, S. M. A., & Rahman, S. C. 2020. Amphibians and reptiles from Lawachara National Park in Bangladesh. Check List, 16: 1239 - get paper here
  • Hora, S.L. 1926. Notes on the Lizard in the Indian Museum. II. On the unnamed Collection of Lizards of the Genus Agamidae. Records of the Indian Museum 28 (4): 215-220
  • Huang, Y.-Z. 1980. A new generic record of chinese lizard from Xizang (Tibet) - Ptyctolaemus gularis Peters. Acta Zootaxonomica Sinica 5: 336
  • Islam, Mazedul & Prasanta Kumar Saikia 2013. Inventory and Natural History of Lizards in Jeypore Rerserve Forest, Assam. Reptile Rap (15): 16-26 - get paper here
  • Lalremsanga, H.T., L. Khawlhring & Lalrotluanga 2010. Three additional lizard (Squamata: Sauria) records for Mizoram, India. Journal of Threatened Taxa 2(2): 718-720 - get paper here
  • Lenz, Norbert 2012. Von Schmetterlingen und Donnerdrachen - Natur und Kultur in Bhutan. Karlsruher Naturhefte 4, Naturkundemuseum Karlsruhe, 124 pp.
  • Mahananda, P., S.N. Jelil, S.C. Bohra, N. Mahanta, R.B. Saikia & J. Purkayastha 2023. Terrestrial vertebrate and butterfly diversity of Garbhanga Landscape, Assam, India. Journal of Threatened Taxa 15(4): 23029–23046 - get paper here
  • Mahony, Stephen; Md. Kamrul Hasan, Md. Mofizul Kabir, Mushfiq Ahmed and Md. Kamal Hossain. 2009. A catalogue of amphibians and reptiles in the collection of Jahangirnagar University, Dhaka, Bangladesh. Hamadryad 34 (1): 80 – 94 - get paper here
  • Malsawmdawngliana, B. Boruah, N.G. Patel, S. Lalronunga, I. Zosangliana, K. Lalhmangaiha & A. Das 2022. An updated checklist of reptiles from Dampa Tiger Reserve, Mizoram, India, with sixteen new distribution records. Journal of Threatened Taxa 14(10): 21946–21960 - get paper here
  • Manthey U 2010. Agamid Lizards of Southern Asia. Draconinae 2 -Leiolepidinae. Edition Chimaira, Terralog 7b, Frankfurt, 168 pp.
  • Peters,W. 1864. Über einige neue Säugethiere (Mormops, Macrotus, Vesperus, Molossus, Capromys), Amphibien (Plathydactylus, Otocryptis, Euprepes, Ungalia, Dromicus, Tropidonotus, Xenodon, Hylodes), und Fische (Sillago, Sebastes, Channa, Myctophum, Carassius, Barbus, Capoëta, Poecilia, Saurenchelys, Leptocephalus). Mber. k. preuss. Akad. Wiss. Berlin [1864]: 381-399 - get paper here
  • Purkayastha J. 2013. An Amateur’s Guide to Reptiles of Assam. EBH Publishers (India) - get paper here
  • Purkayastha, J. 2018. Urban biodiversity: an insight into the terrestrial vertebrate diversity of Guwahati, India. Journal of Threatened Taxa 10(10): 12299–12316; - get paper here
  • Schulte II, J. A., J. V. Vindum, H. Win, T. Thin, K. S. Lwin & A.K. Shein 2004. Phylogenetic relationships of the genus Ptyctolaemus (Squamata: Agamidae), with a description of a new species from Chin Hills of Western Myanmar. Proc. Cal. Acad. Sci., 55(12): 222-247 - get paper here
  • Smith, M.A. 1935. The fauna of British India, including Ceylon and Burma. Reptiles and Amphibia, Vol. II. Sauria. Taylor and Francis, London, 440 pp.
  • Smith, M.A. 1940. The Amphibians and Reptiles obtained by Mr. Ronald Kaulback in Upper Burma. Records of the Indian Museum 42: 465-486 - get paper here
  • Thapa, M. K., Dutta, S., Das, H. J., Pradhan, T. K., Mahanta, D., Tossa, S., ... & Ray, P. C. 2025. Herpetofaunal inventory of the Ranga Reserve Forest, Lakhimpur, Assam, India. Reptiles & Amphibians, 32(1), e21663-e21663
  • Uetz, P.H.; Patel, M.; Gbadamosi, Z.; Nguyen, A.; Shoope, S. 2024. A Reference Database of Reptile Images. Taxonomy 4: 723–732 - get paper here
  • Venugopal, P.D. 2010. An updated and annotated list of Indian lizards (Reptilia: Sauria) based on a review of distribution records and checklists of Indian reptiles. Journal of Threatened Taxa 2 (3): 725-738. - get paper here
  • Wall,F. 1908. Remarks on the Agamoid Lizard (Ptyctolaemus gularis). J. Bombay Nat. Hist. Soc. 18: 505 - get paper here
  • Wang K, Lyu ZT, Wang J, Qi S, Che J 2022. Updated Checklist and Zoogeographic Division of the Reptilian Fauna of Yunnan Province, China. Biodiversity Science 30 (4): 21326, 1–31 - get paper here
  • Wang, Kai; Jinlong Ren, Hongman Chen, Zhitong Lyu, Xianguang Guo Ke Jiang, Jinmin Chen, Jiatang Li, Peng Guo, Yingyong Wang, Jing Che 2020. The updated checklists of amphibians and reptiles of China. Biodiversity Science 28 (2): 189-218 - get paper here
  • Zhao,E. & Adler,K. 1993. Herpetology of China. SSAR, Oxford/Ohio, 1-522
 
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