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Ptyodactylus guttatus HEYDEN, 1827

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Higher TaxaPhyllodactylidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Sinai Fan-fingered Gecko 
SynonymPtyodactylus guttatus HEYDEN 1827
Gecko maculatus SCHINZ 1833 (nom. subst. pro P. guttatus)
Ptyodactylus gecko — GRAY 1845: 151
Ptyodactylus lacazii BOUTAN 1892: 18
Ptyodactylus lacazi — BOUTAN 1893: 444
Ptyodactylus guttatus — KLUGE 1993
Ptyodactylus guttatus — RÖSLER 2000: 107
Ptyodactylus guttatus — LIVIGNI 2013: 401
Ptyodactylus guttatus — BAHA EL DIN 2013
Ptyodactylus guttatus — BAR et al. 2021 
DistributionEgypt, Israel (Gizeh/Beni Hassan, Faroun), Syria, West Jordan, N Saudi Arabia

Type locality: “Gegend von Tor im peträischen Arabien, auch am Sinai”

siphonorhina: Egypt (see map in METALLINOU et al. 2015); Type locality: Abu Roash, near Gizeh.  
Reproductionoviparous 
TypesHolotype: SMF 8369 (formerly 4110d) 
Diagnosis 
CommentSynonymy: after WERNER & SIVAN (1994) who consider the Omani populations of P. hasselquisti as P. guttatus. Previous authors considered P. siphonorhina as a synonym of P. guttatus.

Distribution: Not in Oman (Carranza et al. 2018). See Bar et al. 2021 for a map.

Sexual dimorphism: females are lighter colored and patches are less contrast than in males (Werner, 1993). 
EtymologyThe name guttata is Latin for "spotted" and refers to the numerous dark and light spots and blotches on the back of the species. 
References
  • Al-Quran, S. 2009. The Herpetofauna of the Southern Jordan. American-Eurasian J. Agric. & Environ. Sci., 6 (4): 385-391 [this journal has a dubious record, see http://goo.gl/81SBPv]
  • Anderson, John 1896. A Contribution to the Herpetology of Arabia, with a preliminary list of the reptiles and batrachians of Egypt. London, R. H. Porter, 124 pp. - get paper here
  • Baha el Din, S. 2006. A guide to the reptiles and amphibians of Egypt. The American University in Cairo Press, Cairo. xvi + 359 pp.
  • Bar, A., Haimovitch, G. and Meiri, S. 2021. Field guide to the amphibians and reptiles of Israel. Edition Chimaira, Frankfurt Am Main. ISBN 9783-89973-120-0 - get paper here
  • Bar, Aviad and Guy Haimovitch 2012. A Field Guide to Reptiles and Amphibians of Israel. Pazbar LTD, 246 pp. - get paper here
  • Boutan, L. 1892. Excursion zoologique a la Montagne de Hummoun ul Faroun (Montagne des Bains de Pharaon). Archives de Zoologie Expérimentale et Générale, Paris, 10:1-22
  • Boutan, L. 1893. Mémoire sur les reptiles rapportés de Syrie par le Docteur Théod. Barrois. 1re Partie. Genre Ptyodactyle. Revue Biologique du Nord de la France, 5:1-36 [reprint pagination]. - get paper here
  • Carranza S, Xipell M, Tarroso P, Gardner A, Arnold EN, Robinson MD, et al. 2018. Diversity, distribution and conservation of the terrestrial reptiles of Oman (Sauropsida, Squamata). PLoS One 13 (2): e0190389 - get paper here
  • Disi, A.M.; Modry, D.; Nečas, P. & Rifai, L. 2001. Amphibians and reptiles of the Hashemite Kingdom of Jordan. Edition Chimaira, Frankfurt, 408 pp.
  • Disi, Ahmad M., Zuhair S. Amr and Harald Martens 2004. On a collection of amphibians and reptiles made by J. Klapperich in Jordan. Herpetozoa 16 (3/4):141-150 - get paper here
  • Goldberg, Stephen R. 2011. Ptyodactylus guttatus (Sinai fan-fingered gecko) reproduction. Herpetological Review 42 (3): 433 - get paper here
  • Gray, J. E. 1845. Catalogue of the specimens of lizards in the collection of the British Museum. Trustees of die British Museum/Edward Newman, London: xxvii + 289 pp. - get paper here
  • Handal, Elias N.; Zuhair S. Amr, Mazin B. Qumsiyeh 2016. Some Records of Reptiles from the Palestinian Territories. Russ. J. Herpetol. 23 (4): 261-270 - get paper here
  • Heimes, P. 1987. Beitrag zur Systematik der Fächerfinger (Sauria: Gekkonidae: Ptyodactylus). Salamandra 23 (4): 212-235 - get paper here
  • Heyden, C. H. G. von. 1827. Reptilien. In Rüppell, E. Atlas zu Reise im nördlichen Afrika. l. Zoologie. H. L. Brönner, Frankfurt a. M., pp. 1-24 - get paper here
  • Ibrahim, Adel A. 2013. The Herpetology of the Suez Canal Zone, Egypt. Vertebrate Zoology 63 (1): 87–110 - get paper here
  • Jablonski, D., Sion, G., Bursey, C.R. & Goldberg, S.R. 2017. Ptyodactylus guttatus (Sinai Fan-fingered Gecko) Endoparasite. Herpetological Review 48 (3): 659. - get paper here
  • Jestrzemski, D. 2019. Reptilien in Israel - unterwegs in der Negev-Wüste. Elaphe 2019 (3): 40-49
  • Johnston, G., and A. Bouskila 2007. Sexual dimorphism and ecology of the gecko, Ptyodactylus guttatus. Journal of Herpetology 41: 506–513 - get paper here
  • LiVigni, F. (ed.) 2013. A Life for Reptiles and Amphibians, Volume 1. Chimaira, Frankfurt, 495 pp. - get paper here
  • Meiri, Shai; Amos Belmaker, Daniel Berkowic, Kesem Kazes, Erez Maza, Guy Bar-Oz and Roi Dor 2019. A checklist of Israeli land vertebrates. Israel Journal of Ecology and Evolution - get paper here
  • Metallinou, Margarita; Jan Cervenka, Pierre-Andre Crochet, Lukas Kratochvil, Thomas Wilms, Philippe Geniez, Mohammed Y. Shobrak, Jose C. Brito, Salvador Carranza 2015. Species on the rocks: Systematics and biogeography of the rock-dwelling Ptyodactylus geckos (Squamata: Phyllodactylidae) in North Africa and Arabia. Molecular Phylogenetics and Evolution 85: 208-220 - get paper here
  • Milto, Konstantin D. 2017. New Records of Reptiles on the Red Sea Coast, Egypt, with Notes on Zoogeography. Russ. J. Herpetol. 24 (1): 11-21 - get paper here
  • Müller, J. 1996. Angaben über einige Amphibien - und Reptilienfunde in Jordanien. Sauria 18 (3): 3-10 - get paper here
  • Pola, L., Hejduk, V., Winkelhöfer, T., Zíka, A., Baker, M. A. A., & Amr, Z. S. 2020. Recent Observations on Amphibians and Reptiles in the Hashemite Kingdom of Jordan. Jordan Journal of Natural History, 7: 11-29 - get paper here
  • Qumsiyeh, Mazin 2016. Fauna of Wadi Al-Quff Protected Area: Amphibians, Reptiles and Mammals. Jordan Journal of Natural History (3): 70-79 - get paper here
  • ROCHA, SARA; HERBERT RÖSLER, PHILIP-SEBASTIAN GEHRING, FRANK GLAW, DAVID POSADA, D. JAMES HARRIS & MIGUEL VENCES 2010. Phylogenetic systematics of day geckos, genus Phelsuma, based on molecular and morphological data (Squamata: Gekkonidae). Zootaxa 2429: 1–28 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Schinz,K.R. 1833. Naturgeschichte und Abbildungen der Reptilien. Wiedmann, Leipzig, 240 pp. - get paper here
  • Schlüter, U. 2007. Zur Taxonomie der ägyptischen Fächerfingergeckos. Reptilia (Münster) 12 (66): 10-11 - get paper here
  • Sindaco R., Fedrighini, N. & VEnchi, A. 1995. Contribution to the herpetology of Jordan. Boll. Mus. Reg. Sci. Nat. Torino 13 (1): 389-405
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Sindaco, Roberto; Riccardo Nincheri, Benedetto Lanza 2014. Catalogue of Arabian reptiles in the collections of the “La Specola” Museum, Florence. Scripta Herpetologica. Studies on Amphibians and Reptiles in honour of Benedetto Lanza: pp. 137-164 - get paper here
  • Spawls, Stephen; Tomáš Mazuch & Abubakr Mohammad 2023. Handbook of Amphibians and Reptiles of North-east Africa. Bloomsbury, 640 pp. - get paper here
  • Werner Y L. Sivan N. 1996. Systematics and zoogeography of Ptyodactylus (Reptilia: Sauria: Gekkonidae) in the levant: 3, experimental and natural hybrids of P. guttatus and P. puiseuxi. ISRAEL JOURNAL OF ZOOLOGY. 1996. vol. 42, no. 2, pp. 185-202.
  • Werner, Y. L. 1965. Über die israelischen Geckos der Gattung Ptyodactylus und ihre Biologie. Salamandra 1 (1-2): 15-25 - get paper here
  • Werner, Yehudah L. 2008. Tales of Tails. Gekko 5 (2): 6-18
  • Werner, Yehudah L.;Sivan, Naomi 1993. Systematics and zoogeography of Ptyodactylus (Reptilia: Sauria: Gekkonidae) in the Levant:1. Biometry of three species in Israel. Revista Española de Herpetología 7: 47-64
  • Werner, Yehudah L.;Sivan, Naomi 1994. Systematics and zoogeography of Ptyodactylus (Reptilia: Sauria: Gekkonidae) in the levant : 2, taxonomy, with a review of ecology and zoogeography. Revista Española de Herpetologia 8: 105-122
 
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