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Sauresia habichii WEINLAND, 1863

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Higher TaxaDiploglossidae, Diploglossa, Anguimorpha, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Stout Four-toed Forest Lizard, Habich’s Galliwasp 
SynonymEmbryopus Habichii WEINLAND 1863: 135
Sauresia habichii — SCHOOLS & HEDGES 2024: 234
Sauresia sepsoides — SCHWARTZ & HENDERSON 1988:162 (part)
Sauresia sepsoides — SCHWARTZ & HENDERSON 1991:463 (part)
Celestus sepsoides — HEDGES et al., 2019:17 (part)
Sauresia sepsoides — SCHOOLS & HEDGES 2021:231 (part)
Sauresia sepsoides — LANDESTOY et al. 2022: 205 (part) 
DistributionHaiti

Type locality: Jérémie, Grand’Anse department, Haiti  
Reproductionovoviviparous 
TypesHolotype: ZMB 1310, Collected by D. F. Weinland in 1857–58.
Other Material examined (n=11). HAITI. Grand’Anse. ANSP 38681–83, an adult from 8.0 km S of Marche Leon, collected by S. Blair Hedges, Manuel Leal, Richard Thomas, and Nicholas Plummer on 28 May 1991 (18.51678, -74.08311; 435 m); KU 227597–99, ca 3 km (airline) Sw Corail, 20 March 1966 (18.54806, -73.91232); KU 227802–03 (ca 7.5 km (airline) SSE Roseau, ca 2 km w La Bastille on 20 March 1966 (18.35407, -73.9395); MCZ R-74595, R-74598, David O. Hill, Trou-Bois on Jeremie Road, 1 January 1963; ZMB 1301, David Friedrich Weinland, Jeremie, 1857–1858. 
DiagnosisDiagnosis (n=12). Sauresia habichi has (1) a dorsal pattern of absent/lineate, (2) head markings present, (3) markings in the longitudinal paramedian area absent/present, (4) dots arranged in bars in the lateral band absent, (5) a maximum SVL of 51.9–63.3 mm, (6) ventral scale rows, 122–125, (7) midbody scale rows, 38–41, (8) total lamellae on one hand, 18–19, (9) total strigae on ten scales, 89–113, (10) relative length of all digits on one hindlimb, 11.2–13.1 %, (11) relative distance between the angled subocular and mouth, 0.000594–0.201 %, (12) relative eye length, 2.22– 2.95 %, (13) relative forelimb length, 9.84–13.1 %, (14) relative ear width, 0.364–0.996 %, (15) relative rostral height, 1.83–2.42 %, (16) relative head length, 13.3–15.0 %, (17) relative mental width, 1.49–1.82 %, (18) relative postmental width, 2.11–2.34 %, (19) relative cloacal width, 5.86–6.68 %, (20) relative prefrontal width, 3.39–3.96 %, (21) relative largest supraocular width, 1.96–2.23 %, (22) relative longest finger length, 2.26–2.71 %, (23) relative distance between the ear and eye, 6.64–7.30 %, (24) relative head width, 77.7–78.6 %, (25) relative frontal width, 71.8–77.5 %, (26) relative nasal height, 0.890–1.11 %, (27) relative angled subocular height, 0.909–1.42 %, (28) relative distance between the eye and naris, 3.04–3.59 %, (29) relative canthal iii length, 1.10–1.41 %, (30) relative angled subocular width, 1.67–2.03 %, and (31) relative nasal length, 0.942–1.12 %. The species stem time is 1.19 Ma and the species crown time is 0.00 Ma (Fig. 4).
we distinguish Sauresia habichi from the other species of Sauresia based on a complex of traits. From S. agramma sp. nov., we distinguish S. habichi by the total lamellae on one hand (18–19 versus 17), the relative distance between angled subocular and mouth (0.000594–0.201 versus 0.217–0.262), the relative head width (77.7–78.6 versus 69.7–74.6), the relative frontal width (71.8–77.5 versus 80.2–81.9), the relative angled subocular width (1.67–2.03 versus 2.07–2.47), and the relative nasal width (0.942–1.12 versus 1.36–1.57). From S. cayemitae sp. nov., we distinguish S. habichi by the ventral scale rows (122–125 versus 101–119), the midbody scale rows (38–41 versus 31–37), and the relative head width (77.7–78.6 versus 68.5–77.1). From S. gracilis sp. nov., we distinguish S. habichi by the head markings (present versus absent), the adult SVL (51.9–63.3 versus 51.1), the ventral scale rows (122–125 versus 103), the midbody scale rows (38–41 versus 36), the relative length of digits on one hindlimb (11.2–13.1 versus 13.8), the relative distance between angled subocular and mouth (0.000594–0.201 versus 0.254), the relative eye length (2.22–2.95 versus 2.97), the relative head length (13.3–15.0 versus 15.7), the relative postmental width (2.11–2.34 versus 2.76), the relative cloacal width (5.86–6.68 versus 7.18), the relative largest supraocular width (1.96–2.23 versus 2.44), the relative longest finger length (2.26–2.71 versus 1.96), the relative distance between the ear and eye (6.64–7.30 versus 7.63), the relative head width (77.7–78.6 versus 67.6), the relative frontal width (71.8–77.5 versus 82.9), and the relative width of canthal iii (1.10–1.41 versus 1.82). From S. manicula sp. nov., we distinguish S. habichi by the total lamellae on one hand (18–19 versus 15–17), the total strigae on ten scales (89–113 versus 119–150), the relative longest finger length (2.26–2.71 versus 2.20–2.21), the relative frontal width (71.8–77.5 versus 81.5), and the relative nasal width (0.942–1.12 versus 1.19–1.29). From S. pangnolae sp. nov., we distinguish S. habichi by the ventral scale rows (122–125 versus 104–119), the relative angled subocular width (1.67–2.03 versus 2.16–2.39), and the relative nasal width (0.942–1.12 versus 1.38–1.62). From S. sepsoides, we distinguish S. habichi by the ventral scale rows (122–125 versus 104–119), the midbody scale rows (38–41 versus 34–37), and the relative nasal width (0.942–1.12 versus 1.16–1.60). From S. synoria sp. nov., we distinguish S. habichi by the midbody scale rows (38–41 versus 34–36), the relative prefrontal width (3.39–3.96 versus 3.01–3.30), the relative largest supraocular width (1.96–2.23 versus 1.57–1.91), the relative head width (77.7–78.6 versus 70.8–76.0), the relative frontal width (71.8–77.5 versus 82.0–94.4), and the relative nasal width (0.942–1.12 versus 1.13–1.35). 
CommentSynonymy: previsously considered a synonym of S. sepsoides. 
EtymologyNamed after Edward Habich of Boston, a friend of Weinland who accompanied him during his six-month visit to Jérémie, Haiti, in 1857–1858. 
References
  • SCHOOLS, M., & HEDGES, S. B. 2024. A new forest lizard fauna from Caribbean islands (Squamata, Diploglossidae, Celestinae). Zootaxa 5554(1): 1-306 - get paper here
  • Weinland, D. F 1863. Beschreibung und Abbildung von drei neuen Sauriern. (Embryopus Habichii und Amphisbaena innocens von Haiti, und Brachymeles Leuckarti von Neuholland.). Abh. senckenb. naturf. Ges. (Frankfurt) 4: 131-143 [1862] - get paper here
 
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