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Sauresia agramma SCHOOLS & HEDGES, 2024

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Higher TaxaDiploglossidae, Diploglossa, Anguimorpha, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Denfer Four-toed Forest Lizard 
SynonymSauresia agramma SCHOOLS & HEDGES 2024: 223
Sauresia sepsoides — SCHWARTZ & HENDERSON 1991: 463 (part)
Celestus sepsoides — HEDGES et al., 2019:17 (part)
Sauresia sepsoides — SCHOOLS & HEDGES 2021:231 (part)
Sauresia sepsoides — LANDESTOY et al., 2022: 205 (part) 
DistributionHaiti

Type locality: Morne D’Enfer (southwestern edge of plateau), Sud-Est Department, Haiti (18.33005, -72.37095; 1433 m).  
Reproduction 
TypesHolotype. ANSP 38688, an adult female, collected by S. Blair Hedges, Tiffany Cloud, Miguel Landestoy, and Marcos Rodriguez on 21 November 2011.
Paratypes (n=3). HAITI. Sud-Est. ANSP 38685–7, S. Blair Hedges, Tiffany Cloud, Miguel Landestoy, and Marcos Rodriguez, Morne D’Enfer (southwestern edge of plateau), 21 November 2011. 
DiagnosisDiagnosis. Sauresia agramma sp. nov. has (1) a dorsal pattern of absent, (2) head markings absent/present, (3) markings in the longitudinal paramedian area absent/present, (4) dots arranged in bars in the lateral band absent, (5) a maximum SVL of 53.8–69.1 mm, (6) ventral scale rows, 121–127, (7) midbody scale rows, 37–39, (8) total lamellae on one hand, 17, (9) total strigae on ten scales, 103–138, (10) relative length of all digits on one hindlimb, 11.3–14.9 %, (11) relative distance between the angled subocular and mouth, 0.217–0.262 %, (12) relative eye length, 2.30–3.16 %, (13) relative forelimb length, 10.2–13.6 %, (14) relative ear width, 0.576–0.709 %, (15) relative rostral height, 1.92–2.12 %, (16) relative head length, 13.6–15.2 %, (17) relative mental width, 1.61–2.05 %, (18) relative postmental width, 2.30–2.70 %, (19) relative cloacal width, 6.09–6.82 %, (20) relative prefrontal width, 3.02–3.98 %, (21) relative largest supraocular width, 1.87–2.47 %, (22) relative longest finger length, 2.11– 2.86 %, (23) relative distance between the ear and eye, 6.93–7.86 %, (24) relative head width, 69.7–74.6 %, (25) relative frontal width, 80.2–81.9 %, (26) relative nasal height, 0.883–1.02 %, (27) relative angled subocular height, 0.912–1.13 %, (28) relative distance between the eye and naris, 2.98–3.75 %, (29) relative canthal iii length, 1.17– 1.30 %, (30) relative angled subocular width, 2.07–2.47 %, and (31) relative nasal length, 1.36–1.57 %. The species stem time is 1.58 Ma and the species crown time is 0.00 Ma (Fig. 4).
we distinguish Sauresia agramma sp. nov. from the other species of Sauresia based on a complex of traits. From Sauresia cayemitae sp. nov., we distinguish S. agramma sp. nov. by the ventral scale rows (121–127 versus 101–119), and the relative distance between the ear and eye (6.93–7.86 versus 6.02–6.88). From S. gracilis sp. nov., we distinguish S. agramma sp. nov. by the dorsal pattern (absent versus mottled/lineate), the adult SVL (53.8–69.1 versus 51.1), the ventral scale rows (121–127 versus 103), the midbody scale rows (37–39 versus 36), the total lamellae on one hand (17 versus 18), the total strigae on ten scales (103–138 versus 100), the relative rostral height (1.92–2.12 versus 1.90), the relative head length (13.6–15.2 versus 15.7), the relative postmental width (2.30–2.70 versus 2.76), the relative cloacal width (6.09–6.82 versus 7.18), the relative longest finger length (2.11–2.86 versus 1.96), the relative head width (69.7–74.6 versus 67.6), the relative frontal width (80.2–81.9 versus 82.9), the relative width of canthal iii (1.17–1.30 versus 1.82), the relative angled subocular width (2.07–2.47 versus 1.96), and the relative nasal width (1.36–1.57 versus 1.33). From S. habichi, we distinguish S. agramma sp. nov. by the total lamellae on one hand (17 versus 18–19), the relative distance between angled subocular and mouth (0.217–0.262 versus 0.000594–0.201), the relative head width (69.7–74.6 versus 77.7–78.6), the relative frontal width (80.2–81.9 versus 71.8–77.5), the relative angled subocular width (2.07–2.47 versus 1.67–2.03), and the relative nasal width (1.36–1.57 versus 0.942–1.12). From S. manicula sp. nov., we distinguish S. agramma sp. nov. by the relative distance between angled subocular and mouth (0.217–0.262 versus 0.168–0.188), the relative rostral height (1.92–2.12 versus 1.89–1.90), the relative postmental width (2.30–2.70 versus 2.13–2.16), the relative angled subocular width (2.07–2.47 versus 1.81–2.01), and the relative nasal width (1.36–1.57 versus 1.19–1.29). From S. pangnolae sp. nov., we distinguish S. agramma sp. nov. by the ventral scale rows (121–127 versus 104–119). From S. sepsoides, we distinguish S. agramma sp. nov. by the ventral scale rows (121–127 versus 104–119). From S. synoria sp. nov., we distinguish S. agramma sp. nov. by the midbody scale rows (37–39 versus 34–36), the relative frontal width (80.2–81.9 versus 82.0–94.4), and the relative nasal width (1.36–1.57 versus 1.13–1.35). (Schools & Hedges 2024)


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References
  • SCHOOLS, M., & HEDGES, S. B. 2024. A new forest lizard fauna from Caribbean islands (Squamata, Diploglossidae, Celestinae). Zootaxa 5554(1): 1-306 - get paper here
  • Weinland, D. F 1863. Beschreibung und Abbildung von drei neuen Sauriern. (Embryopus Habichii und Amphisbaena innocens von Haiti, und Brachymeles Leuckarti von Neuholland.). Abh. senckenb. naturf. Ges. (Frankfurt) 4: 131-143 [1862] - get paper here
 
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