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Sceloporus merriami STEJNEGER, 1904

IUCN Red List - Sceloporus merriami - Least Concern, LC

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Higher TaxaPhrynosomatidae, Sceloporinae, Iguania, Sauria, Squamata (lizards)
SubspeciesSceloporus merriami annulatus SMITH 1937
Sceloporus merriami australis WILLIAMS, SMITH & CHRAPLIWY 1960
Sceloporus merriami ballingeri LEMOS-ESPINAL, SMITH, AUTH & CHISZAR 2001
Sceloporus merriami longipunctatus OLSON 1973
Sceloporus merriami merriami STEJNEGER 1904
Sceloporus merriami sanojae LEMOS-ESPINAL in SMITH, LEMOS-ESPINAL & CHISZAR 2003
Sceloporus merriami williamsi LEMOS-ESPINAL, CHISZAR & SMITH 2000 
Common NamesE: Canyon Lizard
S: Lagartija de Cañón
annulatus: Big Bend Canyon Lizard
australis: Southeastern Canyon Lizard
ballingeri: Ballinger’s Canyon Lizard
longipunctatus: Presidio Canyon Lizard
merriami: Merriam's Canyon Lizard
sanojae: Sanoja's Canyon Lizard
williamsi: Williams’ Canyon Lizard 
SynonymSceloporus merriami STEJNEGER 1904
Sceloporus merriami merriami — SMITH & TAYLOR 1950: 131
Lysoptychus merriami — LARSEN & TANNER 1975
Sceloporus merriami — CONANT & COLLINS 1991: 109
Sceloporus merriami — LINER 1994
Sceloporus merriami merriami — BELL et al. 2003
Sceloporus merriami merriami — CROTHER et al. 2012
Sceloporus merriami — HEIMES 2022

Sceloporus merriami annulatus SMITH 1937: 83
Sceloporus merriami annulatus — SCHMIDT & OWENS 1944
Sceloporus merriami annulatus SMITH 1946
Sceloporus merriami annulatus — BARTLETT & BARTLETT 1999: 225
Sceloporus merriami annulatus — BELL et al. 2003
Sceloporus merriami annulatus — LINER & CASAS-ANDREU 2008
Sceloporus merriami annulatus — CROTHER et al. 2012

Sceloporus merriami australis WILLIAMS, SMITH & CHRAPLIWY 1960
Sceloporus merriami australis — BELL et al. 2003
Sceloporus merriami australis — LINER & CASAS-ANDREU 2008

Sceloporus merriami ballingeri LEMOS-ESPINAL, SMITH, AUTH & CHISZAR 2001
Sceloporus merriami ballingeri — LINER & CASAS-ANDREU 2008

Sceloporus merriami longipunctatus OLSON 1973
Sceloporus merriami longipunctatus — BARTLETT & BARTLETT 1999: 227
Sceloporus merriami longipunctatus — BELL et al. 2003
Sceloporus merriami longipunctatus — CROTHER et al. 2012

Sceloporus merriami sanojae LEMOS-ESPINAL in SMITH et al. 2003
Sceloporus merriami sanojae — BELL et al. 2003
Sceloporus merriami sanojae — LINER & CASAS-ANDREU 2008

Sceloporus merriami williamsi LEMOS-ESPINAL, CHISZAR & SMITH 2000
Sceloporus merriarni willaimsi [sic] LEMOS-ESPINAL, CHISZAR & SMITH 2000: 93
Sceloporus merriami williamsi — BELL et al. 2003
Sceloporus merriami williamsi — LINER & CASAS-ANDREU 2008 
DistributionUSA (SW Texas), Mexico (Nuevo Léon, Coahuila, Chihuahua)

merriami: Rio Grande and its immediate tributaries from western Brewster County to SE Val Verde County, Texas; Mexico (Chihuahua [HR 32: 276]). Type locality: East Painted Cave, near mouth of the Pecos River, Texas.

annulatus: S/C Brewster County, Texas. Mexico (E Coahuila). Type locality: East slope of the Chisos Mountains, Brewster County, Texas.

australis: Mexico (Coahuila). Type Locality: 15.6 mi (25.1 km) E Cuatro Ciénegas, Coahuila, Mexico.

ballingeri: Mexico (Durango). Type Locality: La Campana, Sierra La Campana, municipio Tlahualilo, Durango (26°7'39.1"N, 103°41'00.0"W).

longipunctatus: USA (W Texas); Type Locality: Closed Canyon, 14.3 rd mi (23.0 km.) SE Redford, Presidio County, Texas.

merriami: W Texas, Mexico (Coahuila)

sanojae: Mexico (Chihuahua). Type Locality: Rancho Peñoles (27°7'49.6"N, 103°48'45.0"W), 1194m, Chihuahua, Mexico.

williamsi: Mexico (Chihuahua). Type Locality: El Fortín, 51 air km W Rio Grande at 29°35’40”N.  
Reproductionoviparous 
TypesHolotype: USNM 33039
Holotype: INHS (= UIMNH) 25058, EHT-HMS A787 [annulatus]
Holotype: INHS (= UIMNH) 43319, collected 9 July 1958; paratype: BYU [australis]
Holotype: UBIPRO 5094, Julio A. Lemos-Espinal collector, 9 June 2000 [ballengeri]
Holotype: USNM 192744, formerly REO 3286, collected 31 May 1971, by R. Earl Olson [longipunctatus]
Holotype: UBIPRO 7456, collected 24 July 2001 by Julio A. Lemos-Espinal [sanojae].
Holotype: INHS (= UIMNH) 52378, taken by K. L. Williams, E. O. Moll, F. Vuilleumier, and J. E. Williams, 31 August 1962 [williamsi]
Holotype: UBIPRO 5094, an adult male, 9 June 2000, Julio A. Lemos-Espinal [ballingeri] 
DiagnosisDiagnosis: Smith 1939: 285


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Comment 
EtymologyNamed in honor of C. Hart Merriam, the originator of the life-zone theory of biotic ditribution, and chief of the Bureau of Biological Survey, predecessor of the U. S. Fish and Wildlife Service.

S. m. annulatus is named after the Latin noun annulus refers to the banded subcaudal surface, such banding being one of the key characteristics separating this subspecies from Sceloporus merriami merriami.

S. m. australis has been named after the Latin adjective australis, "southern," referring to the range of this southernmost subspecies of S. merriami.

S. m. ballingeri honors Dr. Royce E. Ballinger, of the University of Nebraska, for his many contributions to herpetology and for guidance of Julio A. Lemos-Espinal to his doctoral degree.

S. m. longipunctatus: The Latin adjective long, "long," the noun punctum, "spot," and the suffix -atus, "pertaining to", refer to laterally elongate paravertebral spots, diagnostic of the subspecies.

S. m. sanojae: The subspecific matronym honors Susy Sanoja Sarabia, in honor of and gratitude for her long-term support of Julio A. Lemos-Espinal in the field, laboratory and at home.

S. m. williamsi: The patronym honors Kenneth L. Williams (1934-2017), a specialist in the classification of snakes and the herpetology of Mexico and Honduras. In the original description, two spellings of the name appeared: willaimsi and williamsi. BELL et al. (2003) designated the latter as correct. 
References
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